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A rat named Remy, with sharpened senses of taste and smell, dreams of becoming a chef like his human idol, Auguste Gusteau. Remy and his family live in an old woman's attic in Paris. Once they are found, they evacuate and Remy gets separated from his family.

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He notices Alfredo Linguini, a new chef at Guesteau's, attempting to fix a soup he ruined, and Remy jumps in to fix Linguini's mistakes. Linguini discovers that Remy can understand him, and he convinces Remy to help him cook.

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Remy explores the streets of Paris and finds himself at Gusteau's restaurant. Attempting to make his dream reality, he is encouraged by an imaginary Gusteau to never give up.

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Remy controls Linguini's movements like a marionette by pulling on his hair while hiding underneath his toque. Another chef at the resturant named Colette teaches Linguini the basics of cooking and comes to appreciate someone heeding her advice.

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Skinner, Gusteau's former sous-chef and the new owner of the restaurant, finds out that Linguini is Gusteaus biological son, and realizes that Linguini is to be the new owner of the restaurant. Afraid of giving up his position to the previous "garbage boy", he hides the evidence in an attempt to keep his job. Meanwhile, Linguini's cooking improves with the help of Colette, and they develop a romantic relationship with one another.

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Food critic Anton Ego announces that he'll be dining at Gusteau's and Remy creates the dish ratatouille which reminds Ego of his mother's cooking. Ego asks to meet the chef and is stunned upon being introduced to Remy after the restaurant closes for the evening.

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After Linguini takes credit for Remy's cooking, he and Remy have a falling out. As revenge, Remy leads him and his family to raid the restaurant's pantries for food. Linguini arrives to apologize, but upon discovering the raid, he furiously kicks Remy and the other rats out.

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The next day, Ego writes a review titled "France's Finest", describing Remy (without revealing that he is a rat) as "nothing less than the finest chef in France".

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Remy, Linguini, and Colette open a bistro, called La Ratatouille, which Ego frequently visits, and the rat family settles into the bistro's attic as their new home.