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The ocean holds the largest of Earth's biomes. It covers 70% of the planet's surface.

Life in the ocean is diverse. The smallest creatures that call the ocean home are so tiny that they can only be seen with a microscope. The largest creatures are blue whales, which can be as much as 34 meters (110 feet) long. There are many different ways to live in the ocean, too. Some animals travel huge distances through ocean water. Others stay in the same place on the ocean floor their entire lives. Some burrow beneath the sand while others float near the surface.

The ocean is not the same everywhere. There are many different ecosystems within the ocean depending on conditions such as the water temperature, the amount of sunlight that filters through the water, and the amount of nutrients.

Sunlight breaks through the top layer of ocean water. It can make its way as deep as 200 meters (656 feet). Almost all marine life (about 90%) lives within this top, sunlit layer of the ocean. There, phytoplankton, algae, and plants like seagrass make their own food through the process of photosynthesis and are the start of most marine food chains.

The temperature of ocean water varies depending on its location. Water near the polar regions is colder than water near the equator. Water that is deep in the ocean is colder than water that is near the ocean surface. Many animals and other organisms can only survive at certain temperatures. Others are able to survive at a range of temperatures and can live in more places in the ocean.

The following links give an overview of four different environments where life flourishes in the ocean.

Life in the Intertidal Zone - Where the ocean meets the land, hardy animals, plants, and algae make their homes between the low tide and high tide levels.

Life in the Open Ocean – The largest area of the marine ecosystem, the open ocean is home to swimming fish, drifting plankton, and other creatures.

Life in the Shallow Ocean – In the sunlit parts of the shallow ocean, many animals, plants, and algae thrive on the seafloor as fish zoom above.

Life in the Deep Ocean – This cold, dark world offers some very unusual extreme environments where many living things have found ways to survive.

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The ocean holds the largest of Earth's biomes. It covers 70% of the planet's surface. Life in the ocean is diverse. The smallest creatures that call the ocean home are so tiny that they can only be seen with a microscope. The largest creatures are blue whales, which can be as much as 34 meters (110 feet) long.

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In the very cold places of the world, survival isn't easy. The soil is frozen, its top surface thawing only during summer, and no trees can grow. Yet plants and animals that are adapted for the harsh conditions thrive. This biome is called tundra. Most of the world's tundra is found in the north polar region. It is called Arctic tundra. There is a small amount of tundra on parts of Antarctica that are not covered with ice. Plus, tundra is found on high altitude mountains and is called alpine tundra.

Permafrost is the term given to frozen soil. During the winter months, permafrost reaches the surface of the tundra. It is very cold during the winter, with temperatures reaching -60 degrees Fahrenheit (-51 degrees Celsius). Very few animals are active in these harsh conditions.

In the summer time, the tundra changes. The Sun is out almost 24 hours a day, so the tundra starts to warm up. The permafrost melts at the surface, and plant life grows. However, the permafrost only disappears for a few inches below the surface. There isn't enough soil for trees to grow, so only small plants are found in the tundra.

At the same time, a variety of animals come out to feast on the plants. Insects come to feed on the animals, and birds appear to enjoy the insects.

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In the very cold places of the world, survival isn't easy. The soil is frozen, its top surface thawing only during summer, and no trees can grow. Yet plants and animals that are adapted for the harsh conditions thrive. This biome is called tundra. Most of the world's tundra is found in the north polar region.

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Tropical rainforests are home to thousands of species of animals, plants, fungi and microbes. Scientists suspect that there are many species living in rainforests have not yet been found or described.

There are areas of rainforests where plants are densely packed. Areas where sunlight can reach the surface are full of interesting plants. In other areas a canopy, made from the branches and leaves of tall trees, shades the ground below, preventing smaller plants from growing.

Rainforests get their name because they receive a lot of rain - an average of 80 inches (203 cm) a year! Rainforests are found at and near the equator, where it is always warm and muggy. The temperature doesn't change very much during the year.

Learn more about the animals that live in tropical rainforests by exploring the links below.

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Tropical rainforests are home to thousands of species of animals, plants, fungi and microbes. Scientists suspect that there are many species living in rainforests have not yet been found or described. There are areas of rainforests where plants are densely packed. Areas where sunlight can reach the surface are full of interesting plants.

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Over one quarter of the Earth's surface is covered by grasslands. Grasslands are found on every continent except Antarctica, and they make up most of Africa and Asia. There are several types of grassland and each one has its own name. Prairies, plains and savannas are all grasslands.

Grasslands develop where there isn't enough rain for forests but too much rain for deserts. Grasslands are filled with - you guessed it - grass. There are many types of grass, though. Fields of wheat are considered grasslands, even though they are often cultivated by people. Grass is special because it grows underneath the ground. During cold periods the grass can stay dormant until it warms up.

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Over one quarter of the Earth's surface is covered by grasslands. Grasslands are found on every continent except Antarctica, and they make up most of Africa and Asia. There are several types of grassland and each one has its own name. Prairies, plains and savannas are all grasslands.

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Deserts are full of interesting questions. How can anything survive in a place with hardly any water? Why is it so dry to begin with?

You can find at least one desert on every continent except Europe. Each desert is different in some way, but they all have one thing in common. In order for an area of land to be considered a desert, it must receive less than 10 inches of water a year.

Clouds are scarce in deserts. Without clouds, there can't be rain, snow, or any other precipitation. Clouds also shade the land, so without them, the desert gets mighty hot as the Sun beats down during the day. At night, the desert can become very cold, because there isn't moisture in the air to hold onto the heat.

The geology of each desert is unique. Some deserts have sand dunes - great waves of sand weathered from rock, that move over time as wind blows the sediment. Other deserts have no dunes but instead have unique rock formations carved by wind and streams that only flow at times when there is moisture.

Many plants and animals survive in these vast, dry lands. Learn more about life in the desert by exploring the links below.

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Deserts are full of interesting questions. How can anything survive in a place with hardly any water? Why is it so dry to begin with? You can find at least one desert on every continent except Europe. Each desert is different in some way, but they all have one thing in common.

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Location | Weather | Plants | Animals | People | Links

LOCATION: There are two types of rainforest biomes: temperate and tropical rainforests. Temperate rainforests are found along coasts in temperate regions. The largest temperate rainforests are on the Pacific coast in North America, stretching from Alaska to Oregon. Other temperate rainforests are found along the coast of Chile, the United Kingdom, Norway, Japan, New Zealand, and S. Australia. Tropical rainforests are generally found between 30°N and 30°S latitudes, covering 6 - 7% of the Earth's land surface. Tropical rainforests can be found around the world: In Central and South America; in Western Africa, eastern Madagascar, and the Zaire basin; and in Indo-Malaysia along the west coast of India, Assam, Southeast Asia, New Guinea, and Queensland, Australia.

WEATHER: Rainforests are important because they help maintain global weather patterns and rain. Water that evaporates from trees falls in other areas as rain.

Tropical rainforests are lush and warm all year long! Temperatures don't even change much between night and day. The average temperature in tropical rainforests ranges from 70 to 85°F (21 to 30°C). The environment is pretty wet in tropical rainforests, maintaining a high humidity of 77% to 88% year-round. The yearly rainfall ranges from 80 to 400 inches (200 to 1000 cm), and it can rain hard. It can downpour as much as 2 inches (5 cm) in an hour!

Temperate rainforests are also wet, but not as rainy as tropical rainforests. It rains about from 60 - 200 inches (150 - 500 cm) each year, while the other moisture comes from the coastal fog that lingers on the trees. The fog provides about 7 - 12 inches (18 - 30°C) of rain each year. Temperate rainforests are a lot cooler than tropical rainforests, but the temperatures are still mild. They often have two distinct seasons: one long wet winter, and a short drier summer.

PLANTS: One type of plant often found in a rainforest is an epiphyte. Epiphytes are plants that live on the surface of other plants, especially the trunks and branches. They often grow on trees to take advantage of sunlight in the canopy. In temperate rainforests common epiphytes are mosses and ferns, while in tropical rainforests there are many kinds of epiphytes, including orchids and bromeliads. There are more than 20,000 varieties of orchids found in the rainforest.

There are about 10 -2 0 species of trees in temperate rainforests that are mostly coniferous, meaning they have needles. Trees in temperate rainforests can live much longer than humans, some live for up to 1000 years! Tropical rainforests have a bigger variety of trees, hundreds of species in fact! These trees are mostly broadleaf trees and have a shorter lifespan. They usually live for 50 - 100 years.

Most trees in tropical rainforests have thin, smooth bark. They don't need thick bark to keep them from drying out because the rainforest is so wet. Also, smooth bark makes it difficult for other plants, such as epiphytes, to grow on the tree surface. Trees often have buttresses, large branching ridges near the base, for support because their roots are often shallow and they grow tall to reach the sunlight. Prop roots also help support trees in shallow soils. Many plants in the rainforests have adapted leaf shapes that help water drip off the plant to avoid too much moisture, which might make bacteria and fungus grow.

Tropical rainforests are so big that they are divided into four zones. The top layer of the rainforest is called the emergent layer. Giant trees grow here that are much taller than the trees below. The next layer is the canopy. It contains trees standing 60 to 150 feet (18 to 45 meters) tall. Their branches form a canopy, like a big beach umbrella that shades the forest floor. Thick, woody vines are found in the canopy. Over 2,500 species of vines grow in the rainforest. Some vines, called lianas, are sometimes as big around as a person! They climb the trees in the canopy to reach for sunlight. The next layer, the understory, is a dark, cool area below the canopy, but above the ground. The understory is shaded from much of the sunlight by the canopy. The forest floor is the bottom layer of the rainforest. This is the area where fallen, decomposing plants and trees lay on the ground. Many insects live here. Temperate rainforests have all of these zones except the emergent layer. The tallest trees in the temperate rainforest canopy grow to be about 300 feet (90 meters) tall.

ANIMALS: Tropical rainforests are home to half the plant and animal species on Earth. Scientists believe that there is such a great diversity of animals in tropical rainforests because they are one of the oldest ecosystems on earth. Some forests in Southeast Asia have been around for at least 100 million years, ever since dinosaurs roamed the earth. Animals in the tropical rainforest are specially adapted to live in this unique environment. A common characteristic found among mammals, birds, reptiles and amphibians, is an adaptation to living in trees. One example is New World monkeys that have prehensile tails that curl around branches allowing the monkey to hold onto the tree with its tail! Other animals are brightly colored, sharply patterned, have loud vocalizations, and like to eat lots of fruit. Most of the animals in the tropical rainforest live in the canopy. There is so much food available up there that some animals never go down to explore the forest floor! Birds are important to rainforests because they like to eat seeds and fruit. Their droppings grow into new plants and help rainforests to survive. In turn, tropical rainforests are important to birds because they provide winter grounds as migratory destination. Parrots are not the only type of birds you will see in the rainforest. In fact, about 27% of the bird species in the world live in tropical rainforests. Insects make up the largest single group of animals that live in tropical forests. They include bright beautiful butterflies, menacing mosquitoes, camouflaged stick insects, and colossal colonies of ants.

In temperate rainforests you'll find a different set of amazing animals. Most of these animals live on, or near the forest floor where they are protected from the wind and rain by the trees above. Many birds and small mammals, such as chipmunks, like to eat seeds that fall on the forest floor. Lots of insects live in the temperate rainforest. Most of them live in tree bark, decomposing dead plant matter, or mossy areas. Birds and amphibians like to eat these insects. Some mammals in the temperate rainforest include deer and bobcats. Cougars and bobcats are the top predators in this biome.

PEOPLE AND THE RAINFOREST: Tropical rainforests are important because they provide oxygen, take in carbon dioxide, and are a huge source of biodiversity and cultural diversity. However, people also rely on tropical rainforests for food, medicine, timber, travel, and more. Rainforests around the world provide people with food and spices, for example, allspice, vanilla, cacao, cassava, ginger, bananas, black pepper, sugar cane, nutmeg and more. Many people are also moving from crowded cities where they cannot find jobs into the rainforest where they are becoming small-scale farmers. With all of this use, we need to be concerned about the stress we are putting on rainforests. Thirty acres of trees are cut in tropical rainforests every minute! Some scientists estimate that rainforests the size of Pennsylvania are lost each year. There are many causes of deforestation. People are logging for firewood, charcoal, building materials and other uses. Trees are being removed for commercial agriculture, which may cause permanent damage. Converting rainforests to pasture land for cattle ranching has destroyed many rainforests. Mining for gold, bauxite, and other minerals can destroy the land, and make it vulnerable to erosion.

What can you do to help our world's rainforests? One way to start helping is to learn more about the plants, animals and people, as well as the issues surrounding rainforests and deforestation. When you learn about new issues, write letters to governmental officials and the newspaper suggesting ways to help solve the problem. Practice conservation at home. This will help reduce the use of rainforests, as well as ecosystems all over the world. Bauxite is a mineral used to make aluminum. By recycling, you will reduce the need for bauxite, which is mined in tropical rainforests. Pets from other countries may have been taken from the wild or imported illegally from tropical countries. If you buy pets that are captively bred in the United States, you will be sure that they didn't come from the rainforest, or another unique environment. Investigate the source of the wood for your furniture to see where it comes from, and if it was harvested sustainably. All of these actions will help protect rainforests all over the world.
LINKS: Rainforest Education Cool Planet: Tropical Rain Forest Rainforest Species Profiles Explore Africa's Rainforest National Geographic: Rainforests at Night Mongabay.com Geography for Kids: Tropical Rain Forests Tropical Rainforests: What's it Like Where You Live?

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Location | Weather | Plants | Animals | People | Links LOCATION: There are two types of rainforest biomes: temperate and tropical rainforests. Temperate rainforests are found along coasts in temperate regions. The largest temperate rainforests are on the Pacific coast in North America, stretching from Alaska to Oregon.

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Location | Weather | Plants | Animals | People | Games | Links

LOCATION: Savannas are comprised mostly of grasses and a few scattered trees. They cover half the surface of Africa, large areas of Australia, South America, and India. That is a lot of the earth's surface! Savannas can result from climate changes, soil conditions, animal behavior, or agricultural practices. Humans create savannas by burning grasslands and cutting down trees so they can plant crops. Large animals, like elephants, can turn a forest into a savanna by knocking trees down, stripping the bark from the trees, and tramping on tree seedlings.

WEATHER: An important factor in the savanna is climate. The climate is usually warm and temperatures range from 68° to 86°F (20 to 30°C). Savannas exist in areas where there is a 6 - 8 month wet summer season, and a 4 - 6 month dry winter season. The annual rainfall is from 10 - 30 inches (25 - 75 cm) per year. During the dry season, lightning often strikes the ground, igniting the dry grasses that cover the savanna.

PLANTS: The savanna is dominated by grasses such as Rhodes grass, red oats grass, star grass, lemon grass, and some shrubs. Most savanna grass is coarse and grows in patches with interspersed areas of bare ground. You won't see many trees in the savanna because of little rainfall. Occasionally, you'll find individual trees or small groves of trees. These mostly live near streams and ponds. The Acacia tree is an interesting plant in the savanna. It has an umbrella shape, with branches and leaves high off the ground that giraffes like to eat. Baobab trees also live in the savanna. They deal with dry conditions by storing water between the bark and meat of the tree.

ANIMALS: There are many different types of animals that live in the savanna. The species found in savannas vary by the geographic location of the biome. Animals native to African savannas include African elephants, zebras, horses, and giraffes. Many animals in the savanna are herbivores, which means they eat plants, and there is plenty of grass in the savanna. During the rainy months animals thrive in the savanna, but the rainy season is only half the year. During the dry season, surface water from the rain is quickly absorbed into the ground by thirsty soils. The competition for water during the dry season is so intense that most birds and many of the large mammals migrate elsewhere in search of water. Depending on the severity of the drought, the migration may be to a place nearby, or far away. The dry season is often associated with fires. Many insects with short life spans die in these fires, but the birds and larger animals are usually able to fly or run to safety. Although small burrowing animals probably can't outrun the flames, they often survive the fire by digging deep into the ground and remaining there until the flames are gone. Some birds, such as the Fork-tailed Drongos, don't flee the fires; they actually fly to the fires. For these birds fire means dinner. They eat the fleeing or flame-roasted insects.

PEOPLE AND THE SAVANNA: Some environmental concerns with savannas include poaching or hunting, overgrazing, and destruction of land for commercial crops. Many animals in the savanna, such as the rhinoceros and zebra, are endangered and threatened with extinction due to hunting, poaching, and habitat loss. The savanna is often damaged when it is used as pastureland for non-native domestic cattle. Cattle grazing also limits the amount of food available for wildlife.
GAMES: LINKS: Blue Planet Biomes: Savanna Enchanted Learning: Savanna Animal Printouts Smithsonian National Zoological Park: African Savanna for Kids

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Location | Weather | Plants | Animals | People | Games | Links LOCATION: Savannas are comprised mostly of grasses and a few scattered trees. They cover half the surface of Africa, large areas of Australia, South America, and India. That is a lot of the earth's surface!

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LOCATION: Although few animals and plants are adapted to the extremely dry desert life, the desert is a vital biome. The desert is important because it covers about a fifth of the earth's surface! There are both hot and cold deserts. Antarctica is the largest desert in the world, while the Sahara in Africa is the largest of the hot deserts. There are also deserts close to Santa Barbara, such as the Mojave the Colorado Desert which encompass parts of Southern California. In North America, there are four major hot, dry deserts, including the Mojave and the Great Basin. Outside the U.S. hot, dry deserts are found in the Southern Asian realm, South and Central America, Ethiopia and Australia.

Another type of desert is the coastal desert, for example, the Atacama Desert in Chile of South America. And then there are cold deserts. That sounds pretty silly! If deserts are supposed to be hot, how can there possibly be a cold desert? Well these deserts are in places like Antarctica and Greenland where vegetation is sparce, just like the more commonly known hot, dry deserts.

WEATHER: Weather is not the same in all deserts. The seasons in hot and dry deserts are usually very hot during the summer and warm during the rest of the year. During winter these deserts get little rainfall. Rain is often light, or in short concentrated bursts. Most of the time evaporation rates are faster than rainfall rates. Sometimes the rain evaporates before even hitting the ground. This is the reason for the dry characteristic of this type of desert. Coastal deserts are in moderately cool to warm areas. Coastal deserts usually have cool winters followed by fairly long, warm summers. The temperature in the winters is generally 41°F (5&degC) or below. In the summer the weather heats up to between 55° and 75°F (12 and 24&degC). Average rainfall is usually 3 - 5 inches (8 - 13 cm). The Atacama is the Earth's driest desert. In the Atacama 1 millimeter or more of rain falls every 5-20 years. Cold deserts have short, moist and moderately warm summers, and long cold winters like one could expect in Antarctica. The winter temperature ranges from -5°F to -110°F (-20.5 to -79&degC), and in the summer it can be a nice, balmy, 32°F (0&degC). The coldest day recorded in Antarctica was -113°F (-80.5&degC)!

PLANTS: Deserts plants have many adaptations to survive in such a dry environment. They are good at storing and finding water. Some plants have seeds that can stay dormant in the sand for a long time, until there is enough rain for them to grow. In hot deserts, you'll often find Cacti. Cacti are great at storing water. With their waxy coating, water can't escape and their spines protect them from being desert dinner. Their roots are shallow, and widely spread so that any rain can be absorbed immediately! Some other plants you might find in the hot desert are creosote bush, sagebrush, and ocotillo. Coastal deserts house a variety of plants. These plants must adapt to minimal rainfall by having extensive root systems that come up to the surface to absorb any possible rainfall, and go far down to absorb any water saturated in the ground. These plants also have very thick leaves that can absorb and store water whenever it is available. The plants that live in coastal deserts include salt bush, rice grass, black sage and chrysothamnus. Plants can even live in cold deserts, but you won't find as many here as in other types of deserts. Plants in cold deserts include algae, grasses, and plants with spiny thin leaves. Usually these plants grow only in the summer.

ANIMALS: Deserts are a very important biome. No, not because of all the sand! Mostly because of all the plants and animals that call the desert "home". Some animals that live in the hot desert are cold-blooded, like snakes, insects, and lizards. Mammals that live in the desert are usually small, such as the kangaroo rat and kit fox. Sometimes it's hard to survive in the desert. Some mice build their home out of fallen cactus spines to protect themselves from predators like coyotes and hawks. In the Eritrean coastal desert in Djibouti, Africa, animals like gazelles, skinks, geckos and dikdiks roam the desert. Fewer animals live in the cold desert. In Antarctica, most of the animals live near the ocean shore. Because of their ice home, seals, penguins, and other birds rely on fish, squid and other sea creatures for their food.

PEOPLE AND THE DESERT: People have a variety of uses for deserts. The desert is popular for tourism and recreation. Often people visit to see the beautiful sand colors and rock formations. Many people enjoy visiting the desert because it is a quiet place to get away from the business and noise of cities and spend time in the open land. The desert offers many types of recreation, for example, rock climbing, hiking and dirt biking. Mining, grazing, road building, and utility projects take place in the desert. Because of desert sensitivity, it is important that desert uses be managed properly to protect the wildlife and habitat.
LINKS: The Deserts in the American Southwest Deserts Environment Geography for Kids: Deserts Cool Planet: Deserts Biomes of the World: Desert
Back to Biomes Index

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Location | Weather | Plants | Animals | People | Links LOCATION: Although few animals and plants are adapted to the extremely dry desert life, the desert is a vital biome. The desert is important because it covers about a fifth of the earth's surface! There are both hot and cold deserts.

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LOCATION: Coral reefs are generally found in clear, tropical oceans. Coral reefs form in waters from the surface to about 150 feet (45 meters) deep because they need sunlight to survive. The three types of reefs include fringing reefs, barrier reefs, and atolls. Fringing reefs occur along shorelines of continents and islands and are commonly found in Hawaii and the Caribbean. Barrier reefs are found farther offshore than fringing reefs, occurring most often in the Indo-Pacific and Caribbean. Atolls are a series of low coral islands surrounding a central lagoon, frequently found in the Indo-Pacific. The largest reef in the world, the Great Barrier Reef in Australia is longer than 1200 miles (1900 km). That's longer than the distance between Seattle, WA and Los Angeles, CA!

HABITAT: Coral reefs need water that is between 68 - 82°F (20 - 28°C), which is often located along the eastern shores of land. Reefs usually develop in areas that have a lot of wave action because the waves bring in food, nutrients and oxygen to the reef. Waves also prevent sediment from falling on the reef. Reefs need calcium from the water to grow, which is more often available in shallow warm waters.

PLANTS: The sun is the source of energy for the coral reef ecosystem. Plant plankton, called phytoplankton, algae and other plants convert light energy into chemical energy through photosynthesis. As animals eat the plants and other animals, energy is passed through the food chain. Reef building corals work together with microscopic algae, called zooxanthellae, that live in their tissue. The zooxanthellae provide oxygen and food to the coral through photosynthesis. The coral polyp gives the algae a home, and the carbon dioxide it needs through respiration. Besides zooxanthellae, algae and seagrasses are the main types of plants in the coral reef ecosystem. These plants give food and oxygen to the animals that live on the reef.Seagrasses are especially important because they provide shelter for juvenile reef animals like conch and lobster.

ANIMALS: Did you know that there can be as many different types of fish in two acres of coral reef in Southeast Asia as there are species of birds on the entire continent of North America? Shocking, isn't it? Coral reefs only make up about 1% of the ocean floor, but they house nearly 25% of life in the ocean. Animals use coral reefs either as a stopping point, like an oasis, as they travel the deep blue sea, or they live as residents at the reef. The corals themselves are the most abundant animal on the reef. They are tiny organisms called polyps, that attach themselves to the hard reef and live there forever. The reef is like a giant apartment building in New York City and the coral polyps live together in each apartment. Corals are closely related to sea anemones and sea jellies, and use their tentacles for defense and to capture their prey. Corals can be a variety of colors, white, red, pink, green, blue, orange and purple, due to natural pigments and the zooxanthellae in their tissues.

Other animals that live on the coral reef include sea urchins, sponges, sea stars, worms, fish, sharks, rays, lobster, shrimp, octopus, snails and many more. Many of these animals work together as a team like the coral polyps and zooxanthellae. This teamwork is called symbiosis. One example of symbiosis on the reef is the anemonefish and sea anemone. The sea anemone's tentacles provide protection and safety for the fish and their eggs, while the fish protects the anemone from predators, such as butterflyfish. Sometimes anemonefish even remove parasites from their home anemone.

PEOPLE AND CORAL REEFS: Coral reef ecosystems are important for many reasons. They remove and recycle carbon dioxide, which is a gas that contributes to global warming. Reefs protect land from harsh weather by absorbing the impact from strong waves and storms. Reefs provide food, for example, lobster and conch. Coral reefs are also a huge tourist attraction. Coral reefs are a big source of biodiversity. Without the reef, many of these plants and animals would die. Some people think coral reefs may provide important medicines for people. For example, some coral skeletons can be used by humans as a bone substitute in reconstructive bone surgery. Coral reefs are also a useful educational tool. People can learn about biomes and ecosystems, and the interrelationship between organisms and their environment by studying coral reefs.

Coral reefs are being destroyed at an alarming rate. It is estimated that we have already lost 10% of the worlds reefs, and scientists say that in the next 50 years many of the coral reefs on Earth will be gone. This destruction is often connected with human activity: pollution, sewage, erosion, irresponsible fishing, poor tourism practices, and global warming.

There are some simple things that you can do to help coral reefs. Don't put chemicals down your drain or on your lawn. Instead use biodegradable products. Even though you may be far from a coral reef ecosystem, these products end up in the watershed and may eventually pollute waters that support coral. Conserve water. The less water you use, the less runoff and wastewater eventually find their way into our oceans. Visit a coral reef! Many vacation spots have beautiful coral reefs. When you go, hire local guides. This way you'll learn about the reef from the people who know it best. When you visit a coral reef, treat it with care, do not touch or step on the corals. Leave the animals where you found them and do not pick them up and move them. If you have an aquarium, buy fish raised in captivity, not caught in the wild. Also, don't use live rock in your aquarium. Although this invertebrate-encrusted rock is still legally harvested in some places, its can hurt the reef habitat. Join a group that is working to protect coral reefs. Pick up trash when you see it, this way it won't find it's way to the ocean. Learn more about coral reefs! Surf the Internet for information about coral reefs and marine conservation organizations. After all, knowledge is power.
GAMES:
LINKS:
Ocean World - Coral Reefs Coral Realm SeaWorld - Corals and Coral Reefs NOAA National Ocean Service - Corals Discovery Kit Reef Education Network NOAA's Coral Reef Information System
Back to Biomes Index

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Location | Habitat | Plants | Animals | People | Games | Links LOCATION: Coral reefs are generally found in clear, tropical oceans. Coral reefs form in waters from the surface to about 150 feet (45 meters) deep because they need sunlight to survive. The three types of reefs include fringing reefs, barrier reefs, and atolls.

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LOCATION:An estuary is an area where seawater mixes with freshwater. Estuaries can be found along the coast. Each day as the tide rises, saltwater is brought into the estuary. Freshwater comes down the rivers and creeks and mixes with this saltwater. During the dry season there may not be much freshwater flowing down the rivers. This can make the estuary very salty. Also, during the dry season water evaporates out of the estuary making it even more salty. If you have ever tasted the water in the ocean, you know it is very salty. Estuaries can become even saltier! (But don't try tasting the water, not only will it taste bad, but it can be bad for your health.)
PLANTS: Plants found in estuaries need to be adapted to salty conditions. Having too much salt can kill many types of plants. Some plants, like pickle weed, can absorb the salt water and store the salt in special compartments, called vacuoles, in the leaves. This makes the plant taste very salty which may be one of the reasons it is called pickle weed. Can you see the other reason in this picture? Other plants, like salt grass and alkali heath, are able to push the salt out onto the surface of the leaves. The salt crystals can be seen if you look very closely.

ANIMALS: Many types of animals are found in estuaries. In fact, the complex food web found in an estuary helps to support an amazing diversity of animals. The decaying plants are eaten by microorganisms (animals so tiny you need a microscope to see them.) The microorganisms are eaten by small invertebrates (animals without backbones.) Fish, like carp and stickleback, then eat the invertebrates. Many of the fish we eat depend upon estuaries. They are sheltered places where fish can have their young and the young fish can grow before moving out into the open ocean. Without estuaries, the number of fish in our oceans would decrease greatly. Occasionally, water in certain areas of the estuary can dry up and all that is left is a mud flat. It may seem like there is no life in this mud flat at all, but there are all kinds of mussels, shrimp, worms and other invertebrates living in the mud. Often you can see birds probing the mud with their beaks looking for a tasty meal.

There are a large number of birds found in estuaries. The migrating birds such as geese use them to rest and feed. There are also birds who are permanent residents of the estuaries. Many of these birds, such as the Great Blue Heron, and the Great Egret, have long legs that they use for wading in the water. Egrets are excellent at hunting fish. Some wiggle their toes in the sand to attract fish, which they can snap up for a meal. If you look at the different types of birds, you will notice that they all have different lengths of bills. This means that each type of bird is adapted to eating different things. This reduces the amount of direct competition between the birds and allows more types of birds to live in the same area.

People and Estuaries: Everyone wants to live near the ocean, right? Unfortunately, in order to build houses near the ocean, estuaries were drained and filled in with dirt so that houses could be built on top of them. Many of our estuaries were destroyed before people realized their value.

Estuaries are valuable for a number of reasons. Many countries and coastal communities around the world rely mainly on fish as their primary source of protein. Estuaries help maintain fish populations in the ocean, and without them we would have fewer fish to eat. Estuaries are also natural water filters. They filter the river water that flows into them, helping to keep our ocean water clean and safe for everyone to enjoy. Another important service they provide is protection from large storms and hurricanes. Large waves caused by these storms can cause incredible amounts of damage when they come onshore. Estuaries create a natural barrier that absorbs the energy of the waves and prevents them from flooding cities, destroying houses and injuring people.
GAME: Water Life: Where Rivers Meet the Sea LINKS: Estuaries.gov National Estuarine Research Reserve System EPA: Exploring Estuaries EPA: National Estuary Program Discovery of Estuarine Environments Office of Naval Research - Ocean Habitats Restore America's Estuaries Natural History of Nova Scotia - Tidal Marsh
Back to Biomes Index

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LOCATION:An estuary is an area where seawater mixes with freshwater. Estuaries can be found along the coast. Each day as the tide rises, saltwater is brought into the estuary. Freshwater comes down the rivers and creeks and mixes with this saltwater. During the dry season there may not be much freshwater flowing down the rivers.

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Temperature

Precipitation

This biome sees 150 to 250 millimeters (6 to 10 inches) of rain per year.

Vegetation

Almost no trees due to short growing season and permafrost; lichens, mosses, grasses, sedges, shrubs

Location

Regions south of the ice caps of the Arctic and extending across North America, Europe, and Siberia (high mountain tops)

Other

Tundra comes from the Finnish word tunturia, meaning "treeless plain"; it is the coldest of the biomes

Example: Aklavik, Canada
Monthly Temperature and Precipitation from 1970 - 2000

Month

Average Monthly Precipitation (mm)

Average Monthly Temperature (°C)

Download the Data (csv)

Description

The tundra is the coldest of the biomes. It also receives low amounts of precipitation, making the tundra similar to a desert. Tundra is found in the regions just below the ice caps of the Arctic, extending across North America, to Europe, and Siberia in Asia. Much of Alaska and about half of Canada are in the tundra biome. Tundra is also found at the tops of very high mountains elsewhere in the world. Temperatures are frequently extremely cold, but can get warm in the summers.

Tundra winters are long, dark, and cold, with mean temperatures below 0°C for six to 10 months of the year. The temperatures are so cold that there is a layer of permanently frozen ground below the surface, called permafrost. This permafrost is a defining characteristic of the tundra biome. In the tundra summers, the top layer of soil thaws only a few inches down, providing a growing surface for the roots of vegetation.

Precipitation in the tundra totals 150 to 250 mm a year, including melted snow. That's less than most of the world's greatest deserts! Still, the tundra is usually a wet place because the low temperatures cause evaporation of water to be slow. Much of the arctic has rain and fog in the summers, and water gathers in bogs and ponds.

Vegetation in the tundra has adapted to the cold and the short growing season. Mosses, sedges, and lichens are common, while few trees grow in the tundra. The trees that do manage to grow stay close to the ground so they are insulated by snow during the cold winters.

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Temperature Precipitation This biome sees 150 to 250 millimeters (6 to 10 inches) of rain per year.

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climate change, global climate change, global warming, natural hazards, Earth, environment, remote sensing, atmosphere, land processes, oceans, volcanoes, land cover, Earth science data, NASA, environmental processes, Blue Marble, global maps

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Temperature

Precipitation

Grasslands receive around 500 to 900 millimeters (20 - 35 inches) of rain per year.

Vegetation

Grasses (prairie clover, salvia, oats, wheat, barley, coneflowers)

Location

The prairies of the Great Plains of North America, the pampas of South America, the veldt of South Africa, the steppes of Central Eurasia, and surrounding the deserts in Australia

Other

Found on every continent except Antarctica

Example: Austin, Texas
Monthly Temperature and Precipitation from 1970 - 2000

Month

Average Monthly Precipitation (mm)

Average Monthly Temperature (°C)

Download the Data (csv)

Description

Grasslands are generally open and continuous, fairly flat areas of grass. They are often located between temperate forests at high latitudes and deserts at subtropical latitudes. Grasses vary in size from 2.1 m (7 ft) tall with roots extending down into the soil 1.8 m (6 ft), to the short grasses growing to a height of only 20 to 25 cm (8 to 10 in) tall. These short grasses can have roots that extend 1 m (about 3 ft) deep.

The height of grass correlates with the amount of rainfall it receives. Grasslands receive about 500 to 950 mm of rain per year compared to deserts, which receive less than 300 mm and tropical forests, which receive more than 2,000 mm. While temperatures are often extreme in some grasslands, the average temperatures are about -20°C to 30°C. Tropical grasslands have dry and wet seasons that remain warm all the time. Temperate grasslands have cold winters and warm summers with some rain.

The grasses die back to their roots annually and the soil and the sod protect the roots and the new buds from the cold of winter or dry conditions. A few trees may be found in this biome along the streams, but not many due to the lack of rainfall.

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Temperature Precipitation Grasslands receive around 500 to 900 millimeters (20 - 35 inches) of rain per year.

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Temperature

Precipitation

Deserts get about 250 millimeters (10 inches) of rain per year-the least amount of rain of all of the biomes.

Vegetation

Cacti, small bushes, short grasses

Location

Between 15° and 35° latitude (North and South of the equator); examples are Mojave, Sonoran, Chihuahua, and Great Basin (North America); Sahara (Africa); Negev (Middle East); and Gobi (Asia)

Other

Perennials survive for several years by becoming dormant and flourishing when water is available. Annuals are referred to as ephemerals because some can complete an entire life cycle in weeks.

Example: Cairo, Egypt
Monthly Temperature and Precipitation from 1970 - 2000

Month

Average Monthly Precipitation (mm)

Average Monthly Temperature (°C)

Download the Data (csv)

Description

Desert biomes are the driest of all the biomes. In fact, the most important characteristic of a desert is that it receives very little rainfall. Most deserts receive less than 300 mm a year compared to rainforests, which receive over 2,000 mm. That means that the desert only gets 10 percent of the rain that a rainforest gets! The temperature in the desert can change drastically from day to night because the air is so dry that heat escapes rapidly at night. The daytime temperature averages 38°C while in some deserts it can get down to -4°C at night. The temperature also varies greatly depending on the location of the desert.

Since desert conditions are so severe, the plants that live there need to have adaptations to compensate for the lack of water. Some plants, such as cacti, store water in their stems and use it very slowly, while others like bushes conserve water by growing few leaves or by having large root systems to gather water. Some desert plant species have a short life cycle of a few weeks that lasts only during periods of rain.

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Temperature Precipitation Deserts get about 250 millimeters (10 inches) of rain per year-the least amount of rain of all of the biomes. Vegetation Cacti, small bushes, short grasses Location Between 15° and 35° latitude (North and South of the equator); examples are Mojave, Sonoran, Chihuahua, and Great Basin (North America); Sahara (Africa); Negev (Middle East); and Gobi (Asia) Other Perennials survive for several years by becoming dormant and flourishing when water is available.

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The Arctic tundra, vast plains filled with grasses, flowers, mosses and lichen, is located north of the taiga forests in Earth's north polar region. Like all types of tundra, this is a very cold and windy place. There is little precipitation. Most of the precipitation is snow which insulates the small plants and ground dwelling animals from the cold.

Winter in the Arctic tundra is long, dark, and very cold with temperatures as low as -70°C (-94°F) at times. However, during the short summer growing season, temperatures climb above freezing, thawing the top layer of the frozen soil - the permafrost - for a few weeks. The ground becomes saturated with the water that had been frozen all winter in the permafrost. The water forms ponds and marshes that pepper the barren landscape during the short summer. With water and milder temperatures, the Arctic tundra becomes full of life each summer.

There are no trees in the Arctic tundra; however, there are many different species of plants including small shrubs, grasses and sedges, mosses, and flowers. There are also many types of lichens. The plants that can survive in this extreme climate have adaptations that protect them from cold temperatures, such as grouping together and staying short. They can photosynthesize at low temperatures and low light intensities. Some plants even have fine hairs along their stems that keep them warm.

Many animals, like caribou, insects and many birds, migrate into the Arctic tundra from lower latitudes during the Arctic summer. Other animals, such as the Arctic hare, musk ox, and ptarmigan, have adaptations that allow them to survive the fierce conditions of the Arctic tundra year-round. Adaptations to the Arctic tundra include thick fur or feathers for warmth. Some animals have fur or feathers that turn white in color during the winter to camouflage against the white snow, and then become darker during summer. Many animals, especially those that stay in the tundra year-round, have the ability to form thick layers of fat on their bodies during summer when food is plentiful. The fat serves as insulation from the cold, and is converted to energy during the winter when food is scarce. Many of the animals that remain in the arctic tundra during the winter hibernate, which allows them to not eat at all during the winter.

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The Arctic tundra, vast plains filled with grasses, flowers, mosses and lichen, is located north of the taiga forests in Earth's north polar region. Like all types of tundra, this is a very cold and windy place. There is little precipitation. Most of the precipitation is snow which insulates the small plants and ground dwelling animals from the cold.

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Grassland biomes are large, rolling terrains of grasses, flowers and herbs. Latitude, soil and local climates for the most part determine what kinds of plants grow in a particular grassland. A grassland is a region where the average annual precipitation is great enough to support grasses, and in some areas a few trees. The precipitation is so eratic that drought and fire prevent large forests from growing. Grasses can survive fires because they grow from the bottom instead of the top. Their stems can grow again after being burned off. The soil of most grasslands is also too thin and dry for trees to survive.

When the settlers of the United States moved westward, they found that the grasslands, or prairies as they called them, were more than just dry, flat areas. The prairies contained more than 80 species of animals and 300 species of birds, and hundreds of species of plants.

There are two different types of grasslands; tall-grass, which are humid and very wet, and short-grass, which are dry, with hotter summers and colder winters than the tall-grass prairie. The settlers found both on their journey west. When they crossed the Mississippi River they came into some very tall grass, some as high as 11 feet. Here it rained quite often and it was very humid. As they traveled further west and approached the Rocky Mountains, the grass became shorter. There was less rain in the summer and the winters got colder. These were the short-grass prairies.

Grassland biomes can be found in the middle latitudes, in the interiors of continents. They can have either moist continental climates or dry subtropical climates. In Argentina, South America, the grasslands are known as pampas. The climate there is humid and moist. Grasslands in the southern hemisphere tend to get more precipitation than those in the northern hemisphere, and the grass tends to be the tall-grass variety.

There is a large area of grassland that stretch from the Ukraine of Russia all the way to Siberia. This is a very cold and dry climate because there is no nearby ocean to get moisture from. Winds from the arctic aren't blocked by any mountains either. These are known as the Russian and Asian steppes.

In the winter, grassland temperatures can be as low as -40° F, and in the summer it can be as high 70° F. There are two real seasons: a growing season and a dormant season. The growing season is when there is no frost and plants can grow (which lasts from 100 to 175 days). During the dormant (not growing) season nothing can grow because its too cold.

In tropical and subtropical grasslands the length of the growing season is determined by how long the rainy season lasts. But in the temperate grasslands the length of the growing season is determined by temperature. Plants usually start growing when the daily temperature reached about 50° F.

In temperate grasslands the average rainfall per year ranges from 10-30 inches. In tropical and sub-tropical grasslands the average rainfall per year ranges from 25-60 inches per year The amount of rainfall is very important in determining which areas are grasslands because it's hard for trees to compete with grasses in places where the uppers layers of soil are moist during part of the year but where deeper layer of soil are always dry.

The most common types of plant life on the North American prairie are Buffalo Grass, Sunflower, Crazy Weed, Asters, Blazing Stars, Coneflowers, Goldenrods, Clover, and Wild Indigos.

Some common animals in the grasslands are Coyotes, Eagles, Bobcats, the Gray Wolf, Wild Turkey, Fly Catcher, Canadian Geese, Crickets, Dung Beetle, Bison, and Prairie Chicken.

by Sam M. 2000
Bibliography:
Living In The Environment: An Introduction To Environmental Science G. Tyler Miller--20th edition; Wadsworth Publishing, Belmont, California,

Encyclopedia Britannica, 20th Edition

"Temperate Grasslands" http://www.runet.edu/~swoodwar/CLASSES/GEOG235/biomes/tempgrass/tempgras.html, (Oct. 2000).

"Grasslands",
http://www.ucmp.berkeley.edu/glossary/gloss5/biome/grasslan.html, (Oct. 2000).

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Grassland biomes are large, rolling terrains of grasses, flowers and herbs. Latitude, soil and local climates for the most part determine what kinds of plants grow in a particular grassland. A grassland is a region where the average annual precipitation is great enough to support grasses, and in some areas a few trees.

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The tropical rain forest is a forest of tall trees in a region of year-round warmth. An average of 50 to 260 inches (125 to 660 cm.) of rain falls yearly.

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In this report you will learn about Hot and Dry Deserts and Cold Deserts. I hope you enjoy!

A Hot and Dry Desert is, as you can tell from the name, hot and dry. Most Hot and Dry Deserts don't have very many plants. They do have some low down plants though. The only animals they have that can survive have the ability to burrow under ground. This is because they would not be able to live in the hot sun and heat. They only come out in the night when it is a little cooler.

A cold desert is a desert that has snow in the winter instead of just dropping a few degrees in temperature like they would in a Hot and Dry Desert. It never gets warm enough for plants to grow. Just maybe a few grasses and mosses. The animals in Cold Deserts also have to burrow but in this case to keep warm, not cool. That is why you might find some of the same animals here as you would in the Hot and Dry Deserts.

Deserts cover about one fifth of the Earth's land surface. Most Hot and Dry Deserts are near the Tropic of Cancer or the Tropic of Capricorn. Cold Deserts are near the Arctic part of the world.

Hot and Dry Deserts temperature ranges from 20 to 25° C. The extreme maximum temperature for Hot Desert ranges from 43.5 to 49° C. Cold Deserts temperature in winter ranges from -2 to 4° C and in the summer 21 to 26° C a year

The precipitation in Hot and Dry Deserts and the precipitation in Cold Deserts is different. Hot and Dry Deserts usually have very little rainfall and/or concentrated rainfall in short periods between long rainless periods. This averages out to under 15 cm a year. Cold Deserts usually have lots of snow. They also have rain around spring. This averages out to 15 - 26 cm a year.

Hot and Dry Deserts are warm throughout the fall and spring seasons and very hot during the summer. the winters usually have very little if any rainfall. Cold Deserts have quite a bit of snow during winter. The summer and the beginning of the spring are barely warm enough for a few lichens, grasses and mosses to grow.

Hot and Dry Deserts vegetation is very rare. Plants are almost all ground-hugging shrubs and short woody trees. All of the leaves are replete (packed with nutrients). Some examples of these kinds of plant are Turpentine Bush, Prickly Pears, and Brittle Bush. For all of these plants to survive they have to have adaptations. Some of the adaptations in this case are the ability to store water for long periods of time and the ability to stand the hot weather.

Cold Desert's plants are scattered. In areas with little shade,about 10 percent of the ground is covered with plants. In some areas of sagebrush it reaches 85 percent. The height of scrub varies from 15 cm to 122 cm. All plants are either deciduous and more or less contain spiny leaves.

Hot and Dry Deserts animals include small nocturnal (only active at night) carnivores. There are also insects, arachnids, reptiles, and birds. Some examples of these animals are Borrowers, Mourning Wheatears, and Horned Vipers. Cold Deserts have animals like Antelope, Ground Squirrels, Jack Rabbits, and Kangaroo Rats.

Stetson N. 2000
Bibliography:
Lambert, Wayne, "Deserts", The World Book Encyclopedia, 1994 ed.

Critchfield, Howard J., "Deserts", The New Book of Knowledge, 1998 ed.

"Deserts". http://www.ucmp.berkeley.edu/glossary/gloss5/deserts.html, (5/30/00)

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In this report you will learn about Hot and Dry Deserts and Cold Deserts. I hope you enjoy! A Hot and Dry Desert is, as you can tell from the name, hot and dry. Most Hot and Dry Deserts don't have very many plants. They do have some low down plants though.

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A savanna is a rolling grassland scattered with shrubs and isolated trees, which can be found between a tropical rainforest and desert biome. Not enough rain falls on a savanna to support forests. Savannas are also known as tropical grasslands. They are found in a wide band on either side of the equator on the edges of tropical rainforests.

Savannas have warm temperature year round. There are actually two very different seasons in a savanna; a very long dry season (winter), and a very wet season (summer). In the dry season only an average of about 4 inches of rain falls. Between December and February no rain will fall at all. Oddly enough, it is actually a little cooler during this dry season. But don't expect sweater weather; it is still around 70° F.

In the summer there is lots of rain. In Africa the monsoon rains begin in May. An average of 15 to 25 inches of rain falls during this time. It gets hot and very humid during the rainy season. Every day the hot, humid air rises off the ground and collides with cooler air above and turns into rain. In the afternoons on the summer savanna the rains pour down for hours. African savannas have large herds of grazing and browsing hoofed animals. Each animal has a specialized eating habit that reduces compitition for food.

There are several different types of savannas around the world. The savannas we are most familiar with are the East African savannas covered with acacia trees. The Serengeti Plains of Tanzania are some of the most well known. Here animals like lions, zebras, elephants, and giraffes and many types of ungulates(animals with hooves) graze and hunt. Many large grass-eating mammals (herbivores) can survive here because they can move around and eat the plentiful grasses. There are also lots of carnivores (meat eaters) who eat them in turn.

South America also has savannas, but there are very few species that exist only on this savanna. In Brazil, Colombia, and Venezuela, savannas occupy some 2.5 million square kilometers, an area about one-quarter the size of Canada. Animals from the neighboring biomes kind of spill into this savanna. The Llanos of the Orinoco basin of Venezuela and Columbia is flooded annually by the Orinoco River. Plants have adapted to growing for long periods in standing water. The capybara and marsh deer have adapted themselves to a semi-aquatic life.

Brazil's cerrado is an open woodland of short twisted trees. The diversity of animals is very great here, with several plants and animals that don't exist anywhere else on earth.

There is also a savanna in northern Australia. Eucalyptus trees take the place of acacias in the Australian savanna. There are many species of kangaroos in this savanna but not too much diversity of different animals

Plants of the savannas are highly specialized to grow in this environment of long periods of drought. They have long tap roots that can reach the deep water table, thick bark to resist annual fires, trunks that can store water, and leaves that drop of during the winter to conserve water. The grasses have adaptations that discourage animals from grazing on them; some grasses are too sharp or bitter tasting for some animals, but not others, to eat. The side benefit of this is that every species of animal has something to eat. Different species will also eat different parts of the grass. Many grasses grow from the bottom up, so that the growth tissue doesn't get damaged by grazers. Many plants of the savanna also have storage organs like bulbs and corms for making it though the dry season.

Most of the animals on the savanna have long legs or wings to be able to go on long migrations. Many burrow under ground to avoid the heat or raise their young. The savanna is a perfect place for birds of prey like hawks and buzzards. The wide, open plain provides them with a clear view of their prey, hot air updrafts keep them soaring, and there is the occasional tree to rest on or nest in. Animals don't sweat to lose body heat, so they lose it through panting or through large areas of exposed skin, or ears, like those of the elephant.

The savanna has a large range of highly specialized plants and animals. They all depend on the each other to keep the environment in balance. There are over 40 different species of hoofed mammals that live on the savannas of Africa. Up to 16 different species of browsers (those who eat leaves of trees) and grazers can coexist in one area. They do this by having their own food preferences, browsing/grazing at different heights, time of day or year to use a given area, and different places to go during the dry season.

These different herbivores provide a wide range of food for carnivores, like lions, leopards, cheetahs, jackals and hyenas. Each species has its own preference, making it possible to live side by side and not be in competition for food.

In many parts of the savannas of Africa people have started using it to graze their cattle and goats. They don't move around and soon the grasses are completely eaten up. With no vegetation, the savanna turns into a desert. Huge areas of savanna are lost to the Sahara desert every year because of overgrazing and farming.

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A savanna is a rolling grassland scattered with shrubs and isolated trees, which can be found between a tropical rainforest and desert biome. Not enough rain falls on a savanna to support forests. Savannas are also known as tropical grasslands. They are found in a wide band on either side of the equator on the edges of tropical rainforests.

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Did you know that the Arctic Tundra is the world’s youngest biome? It was formed 10,000 years ago. Located at latitudes 55° to 70° North, the tundra is a vast and treeless land which covers about 20% of the Earth’s surface, circumnavigating the North pole. It is usually…

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Deciduous forests can be found in the eastern half of North America, and the middle of Europe. There are many deciduous forests in Asia. Some of the major areas that they are in are southwest Russia, Japan, and eastern China. South America has two big areas of deciduous forests in southern Chile and Middle East coast of Paraguay. There are deciduous forests located in New Zealand, and southeastern Australia also.

The average annual temperature in a deciduous forest is 50° F. The average rainfall is 30 to 60 inches a year.

In deciduous forests there are five different zones. The first zone is the Tree Stratum zone. The Tree Stratum zone contains such trees as oak, beech, maple, chestnut hickory, elm, basswood, linden, walnut, and sweet gum trees. This zone has height ranges between 60 feet and 100 feet.

The small tree and sapling zone is the second zone. This zone has young, and short trees. The third zone is called the shrub zone. Some of the shrubs in this zone are rhododendrons, azaleas, mountain laurel, and huckleberries. The Herb zone is the fourth zone. It contains short plants such as herbal plants. The final zone is the Ground zone. It contains lichen, club mosses, and true mosses.

The deciduous forest has four distinct seasons, spring, summer, autumn, and winter. In the autumn the leaves change color. During the winter months the trees lose their leaves.

The animals adapt to the climate by hibernating in the winter and living off the land in the other three seasons. The animals have adapted to the land by trying the plants in the forest to see if they are good to eat for a good supply of food. Also the trees provide shelter for them. Animal use the trees for food and a water sources. Most of the animals are camouflaged to look like the ground.

The plants have adapted to the forests by leaning toward the sun. Soaking up the nutrients in the ground is also a way of adaptation.

A lot of deciduous forests have lost land to farms and towns. Although people are trying to protect the forests some poachers are trying to kill the animals in the forests. The animals are losing their homes because of people building their homes.

by Connie T. 2001
Bibliography:
"All about nature: Biomes-Habits", http://www.zoomwhales.com/biomes/tempdecid/tempdecid.shtml, 11/21/00

"Temperate Deciduous Forest Biome", http://mbgnet.mobot.org/pfg/diverse/temp/index.htm, 11/4/00

"Temperature Deciduous Forest Biome", http://mbgnet.mobot.org/pfg/diverse/temp/locatns.htm, 11/5/00

"Temperature Deciduous Forest Biome", http://mbgnet.mobot.org/pfg/diverse/temp/whats.htm, 11/20/00

"World Biomes", http://www.blueplanetbiomes.org/world_biomes.htm, 5/1/01"

The word "Deciduous" means "falling off or out at a certain season". That explains why deciduous forest means a forest in which the leaves fall off the trees when the winter comes.

The deciduous forests are located in the temperate zone above the tropical forests and below the coniferous forests. Most of Europe, the eastern half of North America, parts of Japan and Asia were once covered with large deciduous forests. Most of the deciduous forests have now disappeared but many of the trees still grow in deciduous forest biome. The types of trees you can find in these three regions are broad leafed deciduous trees and some of the evergreen species. The trees are more commonly known as ash, oak, lime, beech, birch and northern arrowwood. Also found in this biome are wild flowers such as oxlip, bluebells, painted trillium and primrose. As well as things such as carpet moss, tawny milk-cap mushrooms and lady fern.

The soil is very fertile. In fact, some of the great agricultural regions are found in this biome. That is one of the reasons there aren't a lot of original deciduous forests left in the world. Almost all of the forests in North America are second growth forests but it still has the biggest variety of original plant species. In Europe there are only a few species of original trees left. Most of the forests have been cleared for agriculture. China has been clearing the natural trees for at least 4,000 years and most of the forests are man-made.

There are many types of animals in the deciduous forest ranging from mammals like deer to bugs like mosquitoes. Many of the animals are either nut and acorn feeders, or omnivores. Many of the animals have adapted to forest life. Some of them hibernate during the winter months.

A few common animals found in the deciduous forest are, deer, gray squirrels, mice raccoons, salamanders, snakes, robins, frogs and many types of insects. Some animals migrate south when winter comes.

Most deciduous forests are found in Eastern North America somewhere around 35-48° N, and Europe and Asia around 45-60° N. There are some deciduous regions in the southern hemisphere but their plants and animals are different from those of the northern deciduous forests.

The average temperature is around 50° F (about 10° C). The average rainfall is 30-60 inches (75-150 cm ) per year. You can find all four seasons: winter (cold and frosty), summer (hot and humid), fall (cool and breezy), and spring (warm and breezy). There is about a 6 month growing season.

2000
Bibliography:
"Deciduous Forest". World Book Encyclopedia. 1994 ed.

"Deciduous", Websters New World Dectionary, 1994 ed.

"Temperate Deciduous Forest", http://www.coff.edu/ete/modules/msese/earthsysflr/clymo5.html (June 4, 2000).

"Mid-Latitude Deciduous Forest", http://wbgnet.mobot/sets/temp/index.htm (June 4, 2000).

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Deciduous forests can be found in the eastern half of North America, and the middle of Europe. There are many deciduous forests in Asia. Some of the major areas that they are in are southwest Russia, Japan, and eastern China. South America has two big areas of deciduous forests in southern Chile and Middle East coast of Paraguay.

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Location | Weather | Plants | Animals | People | Games | Links

LOCATION: Most temperate, deciduous (leaf-shedding) forests are located in the eastern United States, Canada, Europe, China, Japan, and parts of Russia. Deciduous forests are broken up into five zones. The firstzone is the tree stratum zone. It is the tallest zone and trees here range from 60 to 100 feet (18 to 30 meters) tall. Maple, elm, and oak trees are just some examples of trees found in this zone. The second zone is the small tree and sapling zone. Younger, shorter trees characterize this zone. The shrub zone is the third zone. Shrubs include mountain laurel, huckleberries, and many others. The fourth zone is the herb zone, and contains short herbal plants, like ferns. The Ground zone is the final zone where plants grow directly near the ground. Some plants that grow here are lichens and mosses.

WEATHER: This biome has four changing seasons including winter, spring, summer, and fall. These seasons happen because of the tilt of the Earth's axis. Throughout the year, rays from the sun hit different parts of the world more directly than others, causing varying temperatures, or seasons. If the Earth were not tilted on an axis, temperatures around the globe would always be the same. Temperate deciduous forests also have quite a wet environment. Following rainforests, temperate deciduous forests are the second-rainiest biome. The average yearly precipitation is 30 - 60 inches (75 - 150 cm). This precipitation falls throughout the year, but in the winter it falls as snow. The average temperature in temperate deciduous forests is 50°F (10°C). Summers are mild, and average about 70°F (21°C), while winter temperatures are often well below freezing.

PLANTS: Trees and plants in deciduous forests have special adaptations to survive in this biome. Deciduous trees are trees with leaves rather than pine needles, and they dominate temperate forests. As the seasons change each year, so do the leaves. Each year deciduous trees lose their leaves, and grow them back. In the summer their broad green leaves capture sunlight and help the trees make food through photosynthesis. As temperatures cool in the fall, the chlorophyll (green pigment in leaves) breaks down, causing the beautiful red, yellow and orange leaf colors of fall. In the cold winter, deciduous trees and plants go into dormancy, kind of like sleep. It is too cold for them to protect their leaves from the damage of freezing in the winter, so they simply lse them and seal up the places where the leaves attach to the branch. The warmer spring days signal to the trees that they can grow new leaves again, and restart the cycle.

ANIMALS: Animals in temperate deciduous forests have to adapt to changing seasons. They must be able to cope with cold winters and hot summers. Some animals hibernate or migrate during the winter to escape the cold. Animals who do not hibernate or migrate must have special adaptations to deal with higher exposure to predators in the winter. When leaves fall, there is less cover for animals in this biome to hide from predators.

The black bear is an animal that is well adapted for the temperate deciduous forest biome. It has a heavy coat made of many layers of fur to deal with the winter cold. Black bears have long claws that help them to climb trees.This is an essential adaptation because black bears often live in hollowed trees. Black bears are omnivores, so they eat plants and animals. Most of their diet is composed of plant material, so their long claws are useful to get their food from trees and shrubs. They also hibernate to avoid having to find food in the snowy, frozen winter.

PEOPLE AND THE TEMPERATE DECIDUOUS FOREST: Temperate forests are very important to people as they provide enjoyment as well as many resources including food, timber, and oxygen for us to breathe. However, we are also the cause of some major threats to this biome, one of which is acid rain. Acid rain caused by industrial and vehicle emissions damages the leaves of trees, and causes them to produce smaller and fewer seeds. It also reduces the trees' resistance to disease, pests, and frost. Clear cutting of forests is also a threat to this biome. Trees are cut for timber and land cleared for agriculture. Another problem associated with deciduous forests is the introduction of non-native plant and animal species because it upsets the balance of the forest ecosystem. Non-natives may compete for food and habitat space, possibly threatening the native species.

Although these threats may be worrisome, there are many things that you can do to help protect this unique biome. First of all, you can recycle. Trees are used to make the paper for paper bags, newspapers, printer paper, and many more products we use each day. If you recycle used paper, and make the effort to buy recycled paper, you will be reducing the need to cut down more trees. Also be sure to use both sides of the paper that you write on before you recycle it. Use cloth products instead of paper products, like napkins, towels, plates, and cups. These products can be washed and used again, which helps to save trees. Drive less, and carpool when possible. Car exhaust is one of the main contributors to acid rain. Walk or ride your bike to help keep our environment clean. If you are buying furniture, lumber, or any other wood product, look for the Forest Stewardship Council (FSC) label. This label indicates that the trees were grown in a well-managed forest. Learn more about forests. By reading, searching the Internet, and visiting temperate deciduous forests, you can learn lots of cool things about this biome. You can also teach other people about what you have learned. If we want to continue to enjoy temperate deciduous forests, the products that come from them, and protect the unique habitats within them, we must be sure to do our best to take care of this important biome.
GAMES: LINKS: Geography for Kids: Deciduous Forest Blue Planet Biomes: Deciduous Forest Temperate Deciduous Forest Ecokids Forests Shades of Green: Earth's Forests
Back to Biomes Index

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Location | Weather | Plants | Animals | People | Games | Links LOCATION: Most temperate, deciduous (leaf-shedding) forests are located in the eastern United States, Canada, Europe, China, Japan, and parts of Russia. Deciduous forests are broken up into five zones.The firstzone is the tree stratum zone.

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Temperature

Precipitation

On average, this biome receives 750 to 1,500 millimeters (30 to 59 inches) of rain per year.

Vegetation

Broadleaf trees (oaks, maples, beeches), shrubs, perennial herbs, and mosses

Location

Eastern United States, Canada, Europe, China, and Japan

Other

Temperate deciduous forests are most notable because they go through four seasons: Winter, Spring, Summer, and Fall. Leaves change color (or senesce) in autumn, fall off in the winter, and grow back in the spring; this adaptation allows plants to survive cold winters.

Example: Madrid, Spain
Monthly Temperature and Precipitation from 1970 - 2000

Month

Average Monthly Precipitation (mm)

Average Monthly Temperature (°C)

Download the Data (csv)

Description

Temperate deciduous forests are located in the mid-latitude areas which means that they are found between the polar regions and the tropics. The deciduous forest regions are exposed to warm and cold air masses, which cause this area to have four seasons. The temperature varies widely from season to season with cold winters and hot, wet summers. The average yearly temperature is about 10°C. The areas in which deciduous forests are located get about 750 to 1,500 mm of precipitation spread fairly evenly throughout the year.

During the fall, trees change color and then lose their leaves. This is in preparation for the winter season. Because it gets so cold, the trees have adapted to the winter by going into a period of dormancy or sleep. They also have thick bark to protect them from the cold weather. Trees flower and grow during the spring and summer growing season.

Many different kinds of trees, shrubs, and herbs grow in deciduous forests. Most of the trees are broadleaf trees such as oak, maple, beech, hickory and chestnut. There are also several different kinds of plants like mountain laurel, azaleas and mosses that live on the shady forest floor where only small amounts of sunlight get through.

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Temperature Precipitation On average, this biome receives 750 to 1,500 millimeters (30 to 59 inches) of rain per year. Vegetation Broadleaf trees (oaks, maples, beeches), shrubs, perennial herbs, and mosses Location Eastern United States, Canada, Europe, China, and Japan Other Temperate deciduous forests are most notable because they go through four seasons: Winter, Spring, Summer, and Fall.

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Introduction: The tropical rainforest is earth's most complex biome in terms of both structure and species diversity. It occurs under optimal growing conditions: abundant precipitation and year round warmth. There is no annual rhythm to the forest; rather each species has evolved its own flowering and fruiting seasons. Sunlight is a major limiting factor. A variety of strategies have been successful in the struggle to reach light or to adapt to the low intensity of light beneath the canopy.

Climate: (Koeppen's Af and Am climate types.) Mean monthly temperatures are above 64° F; precipitation is often in excess of 100 inches a year. There is usually a brief season of reduced precipitation. In monsoonal areas, there is a real dry season, but that is more than compensated for with abundant precipitation the rest of the year.

Vegetation: A vertical stratification of three layer of trees is apparent. These layers have been identified as A, B, and C layers:

A layer: the emergents. Widely spaced trees 100 to 120 feet tall and with umbrella-shaped canopies extend above the general canopy of the forest. Since they must contend with drying winds, they tend to have small leaves and some species are deciduous during the brief dry season.
B layer: a closed canopy of 80 foot trees. Light is readily available at the top of this layer, but greatly reduced below it.
C layer: a closed canopy of 60 foot trees. There is little air movement in this zone and consequently humidity is constantly high.
Shrub/sapling layer: Less than 3 percent of the light intercepted at the top of the forest canopy passes to this layer. Arrested growth is characteristic of young trees capable of a rapid surge of growth when a gap in canopy above them opens.
Ground layer: sparse plant growth. Less than 1 percent of the light that strikes the top of the forest penetrates to the forest floor. In such darkness few green plants grow. Moisture is also reduced by the canopy above: one third of the precipitation is intercepted before it reaches the ground.

Growthforms: Various growthforms represent strategies to reach sunlight:

Epiphytes: the so-called air plants grow on branches high in the trees, using the limbs merely for support and extracting moisture from the air and trapping the constant leaf-fall and wind-blown dust. Bromeliads (pineapple family) are especially abundant in the neotropics; the orchid family is widely distributed in all three formations of the tropical rainforest. As demonstration of the relative aridity of exposed branches in the high canopy, epiphytic cacti also occur in the Americas.

Lianas: woody vines grow rapidly up the tree trunks when there is a temporary gap in the canopy and flower and fruit in the tree tops of the A and B layers. Many are deciduous.

Climbers: green-stemmed plants such as philodendron that remain in the understory. Many climbers, including the ancestors of the domesticated yams (Africa) and sweet potatoes (South America), store nutrients in roots and tubers.

Stranglers: these plants begin life as epiphytes in the canopy and send their roots downward to the forest floor. The fig family is well represented among stranglers.

Heterotrophs: non-photosynthetic plants can live on the forest floor.

1. Parasites derive their nutrients by tapping into the roots or stems of photosynthetic species. Rafflesia arnoldi, a root parasite of a liana, has the world's largest flower, more than three feet in diameter. It produces an odor similar to rotting flesh to attract pollinating insects.
2. Saprophytes derive their nutrients from decaying organic matter. Some orchids employ this strategy common to fungi and bacteria.

Common characteristics of tropical trees: Tropical species frequently possess one or more of the following attributes not seen in trees of higher latitudes.

Buttresses: many species have broad, woody flanges at the base of the trunk. Originally believed to help support the tree, now it is believed that the buttresses channel stem flow and its dissolved nutrients to the roots.

Large leaves are common among trees of the C layer. Young individuals of trees destined for the B and A layers may also have large trees. When the reach the canopy new leaves will be smaller. The large leaf surface helps intercept light in the sun-dappled lower strata of the forest.

Drip tips facilitate drainage of precipitation off the leaf to promote transpiration. They occur in the lower layers and among the saplings of species of the emergent layer (A layer).

Other characteristics that distinguish tropical species of trees from those of temperate forests include
Exceptionally thin bark, often only 1-2 mm thick. Usually very smooth, although sometimes armed with spines or thorns.
Cauliflory, the development of flowers (and hence fruits) directly from the trunk, rather than at the tips of branches.
Large fleshy fruits attract birds, mammals, and even fish as dispersal agents.

Soils: Oxisols, infertile, deeply weathered and severely leached, have developed on the ancient Gondwanan shields. Rapid bacterial decay prevents the accumulation of humus. The concentration of iron and aluminum oxides by the laterization process gives the oxisols a bright red color and sometimes produces minable deposits (e.g., bauxite). On younger substrates, especially of volcanic origin, tropical soils may be quite fertile.

Subclimaxes: Distinct communities (varzea) develop on floodplains. Jungles may line rivers where sunlight penetrates all layers of the forest. Where forests have long been cleared and laterites have developed to cause season waterlogging of the substrate, tropical grasslands and palm savannas occur.

Fauna: Animal life is highly diverse. Common characteristics found among mammals and birds (and reptiles and amphibians, too) include adaptations to an arboreal life (for example, the prehensile tails of New World monkeys), bright colors and sharp patterns, loud vocalizations, and diets heavy on fruits.

Distribution of biome: The tropical rainforest is generally found between 10° N and 10° S latitude at elevations below 3,000 feet. There are three major, disjunct formations:

Neotropical ( Amazonia into Central America); Atlantic Rainforest (Mata Atlântica)

African (Zaire Basin with an outlier in West Africa; also eastern Madagascar)

Indo- Malaysian (west coast of India, Assam, southeast Asia, New Guinea and Queensland, Australia.

The species composition and even genera and families are distinct in each. They also differ from species of temperate forests. Species diversity is highest in the extensive neotropical forest; second in the highly fragmented Indo-Malaysian formation; and lowest in Africa. Where 5 to a maximum of 30 species of tree share dominance in the Temperate Broadleaf Deciduous Forest, there may be 40 to 100 different species in one hectare of tropical rainforest. Tropical species of both plants and animals often have very restricted distribution areas.

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Introduction: The tropical rainforest is earth's most complex biome in terms of both structure and species diversity. It occurs under optimal growing conditions: abundant precipitation and year round warmth. There is no annual rhythm to the forest; rather each species has evolved its own flowering and fruiting seasons.

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Introduction: Desert areas are rarely devoid of life. Instead, they abound with wonderfully adapted plants and animals that have evolved various mechanisms for tolerating or avoiding the extremes of aridity and temperature that might be encountered in their environment. Deserts develop under four distinct geographic conditions:

Under zones of high atmospheric pressure associated with the subtropics and centered near 30° latitude. Air descending from the upper atmosphere at these latitudes causes evaporation to exceed precipitation. Much of the Sahara and the Australian desert can be associated with this phenomenon.

West coasts of continents between 20° and 30° latitude. In these latitudes, prevailing winds are easterly and prevent moist air from coming onto the west coast. Cold ocean currents also occur in these locations and moisture in the sea air condenses as fog along the shore. Some of the world's driest deserts are located right on the coast; they received most of their limited precipitation from fog. Such fog deserts include Baja California in North America, the western Sahara in northern Africa; the Atacama in South America, and the Namib in southern Africa.

Rainshadows of high mountain ranges. When air masses are forced over mountains and downslope, they warm and their capacity for holding water vapor increases. Evaporation exceeds precipitation and an arid environment or rainshadow is created on the leeward side. Such conditions account for some of the North American deserts (exemplified in Death Valley, CA), the Patagonian desert in Argentina; and the Peruvian desert.

Interiors of continents. Usually in combination with the rainshadow effect, distance from a major source of moist air results in dry climates in the interior of a land mass. The Great Basin desert of the US, the Australian desert, and the Gobi desert of Mongolia can all be explained in large part to their interior positions.

Climate: Arid climates (BWh and BWk) are those which average less than 10 inches of precipitation a year. Potential evaporation exceeds precipitation in the annual water budget. Furthermore, rainfall is highly localized and relatively unpredictable in terms of when it will occur, although usually there are seasons of highest probability for precipitation. Annual variation in total precipitation may also be great. Temperatures are also variable. They may exceed 100° F on summer afternoons, but dip by 20-30 degrees or more at night. Winters are cool to cold: "hot deserts" rarely experience frost; "cold deserts" may have prolonged periods of below freezing temperatures and snowfall.

Vegetation: Shrubs are the dominant growthform of deserts. They may be evergreen or deciduous; typically have small leaves; and frequently have spines or thorns and/or aromatic oils. Shallow but extensive root systems procure rainwater from well beyond the canopy of the shrub whenever it does rain. These are the true xerophytes adapted to tolerate extreme drought. They form an open canopy and, except after rains when annuals may cover the desert floor, the ground between shrubs is bare of vegetative growth.

Water is not entirely lacking in the desert environment and several other growthforms represent strategies to reach water or to store water:

Phreatophytes are plants with long taproots that may extend downward 20 to 30 feet to tap ground water supplies. Especially along intermittent streams or under dunes, underground water may be readily available. Mesquite is a good example here in North America.
One of the world's most unusual phreatophyes isWelwitschia mirabilis of the Namib.

Succulents store water accumulated during rains for use during the intervening dry spells. Different species store water in different parts of the plant; hence we can recognize stem succulents, leaf succulents, root succulents, and fruit succulents. Many plant families have members that evolved succulence. Most prominent among stem succulents in the Americas are the Cactaceae; in Africa succulent euphorbias have evolved shapes and sizes resembling the cacti. The agaves (Liliaceae) are examples of leaf succulents in the Americas; their role is filled by aloes (Liliaceae) in Africa. Most succulents do not tolerate freezing temperatures so they are essentially limited to the hot deserts.

Another growthform adapted to desert conditions is the ephemeral. This is an especially short-lived annual forb that completes its life cycle in two-three weeks. The seeds are encased in a waterproof coating that prevents desiccation for years if necessary. These plants essentially avoid drought by occurring as seeds most of the time.

Perennial forbs with underground bulbs store nutrients and water in undergound tissues and also remain dormant most of the year. They can sprout rapidly after sufficient rains and replenish their undergound stores.

Soils: Calcification is the dominant soil-forming process, if indeed soil forming even occurs. There is poor development of horizons, with accumulation of calcium carbonate at or near the surface. Sparse vegetative cover and tiny leaves results in little humus and soils typically have a light gray color. Aridosols are the dominant soil order.

Fauna: Like the plants, the animals of the desert have evolved an array of strategies for dealing with aridity.

Behavioral adaptations such as being nocturnal or crepuscular, being fossorial, and staying the shade during the heat of day are common.

Morphological adaptations include those noticed by Bergmann, Allen, and Golger. The better to radiate body heat to the environment from warm-blooded animals, body sizes are small and appendages long. Pelage and plumage is light colored to reflect sunlight and help prevent the absorption of heat form the environment.

Rarer, but important, are physiological adaptations such as aestivation (dormancy during summer), the absence of sweat glands, the concentration of urine, localized deposits of fat in tails or humps; and salt glands to secrete salt without loosing fluids.

Reptiles with their waterproof skin, production of uric acid instead of urine, hard-shelled eggs, and ability to gain body heat directly from the sun and to retreat to shade or underground to avoid heat are exceptionally well adapted to drylands and, not surprisingly, diverse there.

Many birds in the North American deserts, so fragmented by mountains offering humid habitats and permanent streams, simply fly to free water and so are not limited by the lack of open water. They maintain breeding seasons like other temperate zone birds synchronized by changing photoperiods. In Australia, where the desert geography is quite different and aridity more pervasive, bird populations synchronize their breeding readiness according to cues of rainfall, however erratic and sporadic that may be.

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Introduction: Desert areas are rarely devoid of life. Instead, they abound with wonderfully adapted plants and animals that have evolved various mechanisms for tolerating or avoiding the extremes of aridity and temperature that might be encountered in their environment. Deserts develop under four distinct geographic conditions: Under zones of high atmospheric pressure associated with the subtropics and centered near 30° latitude.

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Introduction: Temperate grasslands are composed of a rich mix of grasses and forbs and underlain by some of the world's most fertile soils. Since the development of the steel plow most have been converted to agricultural lands.

Climate: Semiarid, continental climates of the middle latitudes (Koeppen's BSk climate type) typically have between 10 and 20 inches of precipitation a year. Much of this falls as snow, serving as reservoir of moisture for the beginning of the growing season. Warm to hot summers are experienced, depending on latitude.

Vegetation: Perennial grasses and perennial forbs [especially Compositae (or Asteraceae, depending on the taxonomic system used) and Leguminosae--the sunflower and pea families, respectively] are dominant growthforms. Two or more strata of grasses (erect grasses and recumbent species) are recognized in the more humid expressions of the biome.

Grasses. Perennial grasses, with their growth buds at or just below the surface, are well-adapted to drought, fire, and cold. The tiller or narrow, upright stem reduces heat-gain in the hot summers; the intricate root systems trap moisture and nutrients. Two basic types are:

Turf- or sod-forming grasses, with rhizomes or underground stems from which new plants spring forth; associated with the more humid grasslands

Bunch grasses, without rhizomes, that reproduce by seed; associated with the drier parts of the biome.

Major regional expressions:
North America: the prairies of the Central Lowlands and High Plains of the US and Canada. The Palouse Prairie of eastern Washington state, the California grasslands, and the desert grasslands of the Southwest are also temperate grasslands.

Eurasia: the steppes from Ukraine eastward through Russia and Mongolia.

South America: the pampas of Argentina, Uruguay, and southeastern Brazil

Africa: the veld in the Republic of South Africa.

Soils: Calcification is the dominant soil-forming process in semiarid regions. Mild leaching, high organic content, and concentration of calcium carbonate in the B horizon typifies the dark brown mollisols developed under the temperate grasslands. When this process works on a loess that itself is rich in calcium, the world's most fertile soils are created, the chernozems (A Russian term meaning black soil). Loess and hence chernozem underlie the eastern prairies of the US, the pampas of South America, and the steppes of Ukraine and Russia.

Fauna: The temperate grassland fauna is very low in diversity, especially in comparison with the tropical grasslands or savannas of Africa. In North America the dominant herbivores are bison Bison bison) and pronghorn (the sole member of the Nearctic endemic family, Antilocapridae). Rodent herbivores include the pocket gopher (another Nearctic endemic), ground squirrels, and the prairie dog. Carnivores include coyote (actually an omnivore), badger, and the federally endangered black-footed ferret, the last two members of the weasel family.

On the Russian steppes the fauna formerly included wisent ( Bison bonasus), tarpan or wild horse, and saiga antelope, among others. Mole rats, fossorial members of one of the two mammal families endemic to the Palearctic, are conspicuous by virtue their many mounds. Polecats and other members of the weasel family are among the larger, extant carnivores.

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Introduction: Temperate grasslands are composed of a rich mix of grasses and forbs and underlain by some of the world's most fertile soils. Since the development of the steel plow most have been converted to agricultural lands. Climate: Semiarid, continental climates of the middle latitudes (Koeppen's BSk climate type) typically have between 10 and 20 inches of precipitation a year.

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Introduction: Tropical savannas or grasslands are associated with the tropical wet and dry climate type (Koeppen's Aw), but they are not generally considered to be a climatic climax. Instead, savannas develop in regions where the climax community should be some form of seasonal forest or woodland, but edaphic conditions or disturbances prevent the establishment of those species of trees associated with the climax community. Seasonal forests of the tropics are also widespread and vary along a latitudinal/moisture gradient between the tropical broadleaf evergreen forest of the equatorial zone and the deserts of the subtropics.

The word savanna stems from an Amerind term for plains which became Hispanicized after the Spanish Conquest.

Vegetation: Savannas are characterized by a continuous cover of perennial grasses, often 3 to 6 feet tall at maturity. They may or may not also have an open canopy of drought-resistant, fire-resistant, or browse-resistant trees, or they may have an open shrub layer. Distinction is made between tree or woodland savanna, park savanna, shrub savanna and grass savanna. Furthermore, savannas may be distinguished according to the dominant taxon in the tree layer: for example, palm savannas, pine savannas, and acacia savannas.

Climate: A tropical wet and dry climate predominates in areas covered by savanna growth. Mean monthly temperatures are at or above 64° F and annual precipitation averages between 30 and 50 inches. For at least five months of the year, during the dry season, less than 4 inches a month are received. The dry season is associated with the low sun period.

Soils: Soils vary according to bedrock and edaphic conditions. In general, however, laterization is the dominant soil-forming process and low fertility oxisols can be expected.
Regional expressions:
East African savannas are typically, perhaps stereotypically, acacia savannas. Many survive in the famous game parks of Kenya and Tanzania, and also those of Zimbabwe, Botswana, South Africa, and Namibia. The savannas are actually a mosaic of communities controlled (and today managed) by fire and grazing pressures.
The famous Serengeti Plains in Tanzania are a grass savanna developed on droughty but nutrient-rich volcanic sands.

The llanos of the Orinoco basin of Venezuela and Colombia are grass savannas maintained by the annual flooding of the Orinoco and Arauca rivers and their tributaries. The long periods of standing water inhibit the growth of most trees.

is an open woodland of short-stature, twisted trees. It is species-rich, second only to the tropical rainforest in plant diversity. There are many endemic species, and several plants have adaptations to tolerate the high aluminum content of soils resulting from laterization on the ancient Gondwanan Shield of South America.

The pine savannas of Belize and Honduras, in Central America, occur on sandy soils.
Savannas as subclimaxes.
Edaphic Subclimaxes:
Waterlogged conditions occur when the A-horizon of lateritic soils is exposed to the atmosphere. Alternating wet and dry seasons and baking by the sun create a brick-hard layer impermeable to water. This usually red hardpan is called a laterite (from the Latin for brick). During the rainy season, there is standing water above the hardpan for several months, preventing the establishment of most tree species. During the dry season, the laterite prevents penetration of roots, also inhibiting the growth of most trees. Several species of palms do tolerate these conditions and, along with grasses, occur above laterites.

Droughty substrates, such as quartz or volcanic sands, also inhibit the growth of most trees. The pine savannas of Central America are examples of savanna vegetation developed on droughty, low-nutrient conditions of quartz sands; the grass savanna of the Serengeti-with its herds of large mammals-is virtually treeless.
Low-nutrient soils. The cerrado of Brazil occupies a broad expanse of the Brazilian Highlands that, were it not for the low-nutrient level of the heavily-leached soils, would be occupied by a seasonal forest.

Fire subclimaxes. Two groups of plants that are pre-adapted to survive fire become dominant in areas where burning is frequent and periodic. Such fires have both natural and human origins. The savannas of South east Asia are generally considered to be man-made.
Palms have the advantage of being monocots: their vascular bundles are scattered throughout the stem so that scorching of the outermost layer of the trunk will not kill the plant. (Dicot trees, on the other hand, have their vascular bundles arrnaged around the outer, living part of their stems where they may be easily destroyed by fire.)
Perennial grasses have underground stems or rhizomes and so their growth nodes are protected by the soil during a ground fire. Trees and shrubs-with renewal buds above the surface-are selected against by fire and the balance tips toward the grasses.

Grazing subclimax. Large mammals such as the elephant open woodlands by debarking the trees and by knocking them over. This opens the woodland to grass invasion and attracts a variety of grazing animals, including zebras, wildebeest, and the diverse antelopes of the Ethiopian province. Grazers will both eat and trample tree seedlings, inhibiting the regrowth of the woodland. Only well-armed species of shrubs and trees can establish themselves in the clearings, leading to thickets of thorny acacias. Protected in the thicket, some acacias and other thorny trees will grow to mature specimens.

Overgrazing: if a grass savanna is overgrazed, patches of bare ground will be created. The grassland will not longer carry a ground fire and invasion by trees becomes possible. The bare ground will suffer from increased evaporation and a dry microhabitat quickly develops. Well-armed, drought-resistant species like the acacias tolerate both grazing and drought, so again an acacia savanna can become established.

Fauna: The world's greatest diversity (over 40 different species) of ungulates (hoofed mammals) is found on the savannas of Africa. The antelopes are especially diverse and including eland, impalas, gazelles oryx, gerenuk, and kudu. Buffalo, wildebeest, plains zebra, rhinos, giraffes, elephants, and warthogs are among other herbivores of the African savanna. Up to sixteen grazing and browsing species may coexist in the same area. They divide the resources spatially and temporally; each having its own food preferences, grazing/browsing height, time of day or year to use a given area, and different dry season refugia.

The species-rich herbivore trophic level supports a diverse set of carnivores, including cats (lions, leopards, cheetahs, servals), dogs (jackals, wild dogs), and hyenas.

Most herbivorous mammals of the open savannas are herd animals, often organized into groups of females and their young with a single dominant male and groups of bachelor males.

In South America a distinct savanna fauna is not well-developed. The capybara, the large semi-aquatic rodent, is associated with the llanos, but is found elsewhere and in other vegetation types as well. Indeed, few if any neotropical mammals are restricted to the savannas. The highest diversity of mammals is found in the dry or seasonal forests. Similarly, most bird species are not restricted to savanna-type habitats.

Termites are especially abundant in the tropical savannas of the world, and their tall termitarias are conspicuous elements of the savanna landscape. These detrivores are important in soil-formation; their termitaria provide shelter for other animals; and they are the beginning of the food chain for anteaters (Neotropical endemics) and aardvarks and pangolins (Ethiopian endemics).

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Introduction: Tropical savannas or grasslands are associated with the tropical wet and dry climate type (Koeppen's Aw), but they are not generally considered to be a climatic climax. Instead, savannas develop in regions where the climax community should be some form of seasonal forest or woodland, but edaphic conditions or disturbances prevent the establishment of those species of trees associated with the climax community.

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Introduction: The Temperate Broadleaf Deciduous Forest (TBDF)-especially in eastern North America, where is remains most intact-is known for the turning of the colors of its leaves to brilliant reds, oranges, and golds in autumn. The shortening days of fall stimulate the plants to withdraw chlorophyll from their leaves, allowing a brief but beautiful display of other pigments before the leaves are shed completely and plants enter an extended period of dormancy.

Climate: Associated with warmer continental and humid subtropical climates (Dfa, Cfa, and-in Europe, Cfb). There is an approximately 6 month growing season. The 20 to 60 inches of precipitation is distributed evenly throughout the year. The non-growing season is due to temperature-induced drought during the cold winters.

Vegetation: Many of the same genera, previously part of an Arcto-Tertiary Geoflora, are common to all three of the disjunct northern hemisphere expressions of this biome. Included among these genera are Quercus (oak), Acer (maple), Fagus (beech), Castanea (chestnut), Carya (hickory), Ulmus (elm), Tilia (basswood or linden), Juglans (walnut), and Liquidamber (sweet gum). Different species of these genera occur on each continent.

Structure and Growthforms: Five layers are recognized:

a tree stratum, 60 -100 feet high, dominated regionally by various combinations of the genera listed above;

a small tree or sapling layer, with not only younger specimens of the tall trees with species limited to this layer such as (in Virginia) Allegheny serviceberry or shadbush, sourwood, dogwood, and redbud;

a shrub layer often with members of the heath family such as rhododendron, azaleas, mountain laurel, and huckleberries;

an herb layer of perennial forbs that bloom primarily in early spring; and

a ground layer of lichens, clubmosses, and true mosses. Lichens and mosses also grow on the trunks of trees.

Lianas such as wild grape, poison ivy, and Virginia creeper climb the trees to flower and fruit high in the forest canopy.

Soils: Brown forest soils (alfisols, in the American soil taxonomy) develop under the TBDF. Broadleaf trees tend to be nutrient-demanding and their leaves bind the major nutrient bases. Thus the litter under this forest is not as acidic as under needleleaf trees and aluminum and iron are not mobilized from the A horizon. The autumn leaf fall provides for an abundant and rich humus which begins to decay rapidly in spring just as the growing season begins. The humus content gives both A and B horizons a brown color. [Until John Deere's invention of the steel plow in the 1800s and the subsequent ability to break the prairie sod, the alfisols were considered the most fertile, most easily worked, and most easily cleared of northern hemisphere temperate zone soils. Many have been under continuous cultivation since the Neolithic.]

Ultisols replace alfisols in the southeastern US, where the older soils of unglaciated regions have been weathered to a much greater degree and are more completely leached than the younger soils to the north. Distinctive red or yellow subsoils have developed under the warmer climate. Ultisols are generally less fertile than alfisols and in the southeast were frequently further degraded under plantation and subsistence agriculture in both the colonial and post-colonial periods.

Subclimaxes: On sandy substrates, pines replace broadleaf species. Hence the New Jersey pine barrens, the pineywoods of the Deep South, and the tall (long-needled) pines of Georgia and other areas of the Atlantic Coastal Plain. On waterlogged sites in more northerly latitudes, bogs develop. In the south one finds instead pine savannas and bald cypress swamps.

Fauna: Characteristic members of the fauna are either mast-eaters (nut and acorn feeders) or omnivores. Mammals show adaptations to an arboreal life; a few hibernate during the winter months.

North American herbivores include white-tail deer, gray squirrel, and chipmunk.

Omnivores include raccoon, opossum, skunk, and black bear.

Carnivores have been largely eliminated through the deliberate effort of humans but should include timber wolves, mountain lions, and bobcats. The coyote, native to the western grasslands and deserts, has recently dispersed east and taken over the niche of its departed cousin, the timber wolf.

Resident bird species also tend to be seed-eaters or omnivores. Many, like the several species of woodpeckers and the chickadees, are cavity-nesters. The loud, conspicuous blue jay is a major agent in the dispersal of oaks onto abandoned farmland and pastures. Migratory species tend to be insectivorous and include many so-called neotropical migrants, including warblers, wrens, thrushes, tanagers, and hummingbirds.

Distribution: The TBDF occurs in three major, disjunct expressions in western and central Europe; eastern Asia, including Korea and Japan; and eastern North America.

In Europe, a species-poor forest reflects widespread extinctions during the Pleistocene. Oaks, beeches, and elms dominate. Most of the forest was cleared for agriculture, with remnants surviving only in some royal hunting preserves.

The TBDF of China is known primarily from the fossil record; intensive agriculture has caused this region to be cleared of natural vegetation for at least 4,000 years. Japan has a largely artificial forest, but in the mountains of Korea the forest is more or less intact and fall foliage is reminiscent of New England's.

Almost all the forests of eastern North America are second growth, but they preserve the world's greatest diversity of TBDF flora and fauna. This is especially true of the unglaciated Appalachian Plateau of eastern Kentucky and Tennessee and western North Carolina and Virginia. The Great Smoky Mountains have been designated a world biosphere reserve.

Southern hemisphere expressions of the biome: Regions of humid subtropical climate occur in the southern hemisphere, but their vegetation and flora differ. Such regions have a mixed (broadleaf and needleleaf) evergreen forest whose biogeographic interest stems from the occurrence of Gondwanan relicts: Araucaria pines (South America and Australia), Podocarpus pines (South America, Africa, and Australia), and the evergreen southern beech Nothofagus (South America, Australia, and New Zealand).

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Introduction: The Temperate Broadleaf Deciduous Forest (TBDF)-especially in eastern North America, where is remains most intact-is known for the turning of the colors of its leaves to brilliant reds, oranges, and golds in autumn.

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Desert The defining characteristic of a desert is that it is dry. Depending on its geographical location, the annual precipitation in a desert varies from half an inch to as much as 15 inches. Rainfall is usually very localized, and although it is frequently seasonal, it is difficult to predict when or where it will occur. At times in the Atacama Desert in Chile, years have passed with no measurable rainfall at all. However, that is not generally the case. Photo © 2000-www.arttoday.com

Deserts can be either hot such as the Australian Desert or cold such as the Gobi Desert. As with all biomes, the desert climate is determined by geographic conditions. Geographic conditions such as location, high atmospheric pressure, and proximity of mountain ranges determine just what type of desert it is.

Deserts may occur along the coast such as the Atacama and Namib deserts or in the interior of continents such as the Great Basin and Australian deserts, which are far from any source of water. Coastal deserts are located on west coasts of continents between 20° to 30° latitude. Prevailing winds blow in an easterly pattern and prevent the moisture from moving onto the land. Semiarid deserts, like the Great Basin Desert, are not only located far from moisture, but are frequently associated with high mountain ranges that produce a rainshadow effect. The rainshadow effect prevents available moisture from reaching the area. The great Gobi Desert of Mongolia has little rainfall because the Himalayan Mountains prevent rainfall from moving into this region.

Because all deserts are dry, they have large daily temperature variations. Temperatures are high during the day because there is very little moisture in the air to block the Sun's rays from reaching Earth. Once the Sun goes down, the heat absorbed during the day quickly escapes back into space. High daytime temperatures and low nighttime temperatures make survival in the desert very difficult.

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Desert The defining characteristic of a desert is that it is dry. Depending on its geographical location, the annual precipitation in a desert varies from half an inch to as much as 15 inches. Rainfall is usually very localized, and although it is frequently seasonal, it is difficult to predict when or where it will occur.

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Tropical Rainforest
The tropical rainforest is a hot, moist biome found near Earth's equator. The world's largest tropical rainforests are in South America, Africa, and Southeast Asia. Tropical rainforests receive from 60 to 160 inches of precipitation that is fairly evenly distributed throughout the year. The combination of constant warmth and abundant moisture makes the tropical rainforest a suitable environment for many plants and animals. Tropical rainforests contain the greatest biodiversity in the world. Over 15 million species of plants and animals live within this biome. Photo © Gayle W. Croft

The hot and humid conditions make tropical rainforests an ideal environment for bacteria and other microorganisms. Because these organisms remain active throughout the year, they quickly decompose matter on the forest floor. In other biomes, such as the deciduous forest, the decomposition of leaf litter adds nutrients to the soil. But in the tropical rainforest, plants grow so fast that they rapidly consume the nutrients from the decomposed leaf litter. As a result, most of the nutrients are contained in the trees and other plants rather than in the soil. Most nutrients that are absorbed into the soil are leached out by the abundant rainfall, which leaves the soil infertile and acidic.

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Tropical Rainforest The tropical rainforest is a hot, moist biome found near Earth's equator. The world's largest tropical rainforests are in South America, Africa, and Southeast Asia. Tropical rainforests receive from 60 to 160 inches of precipitation that is fairly evenly distributed throughout the year.

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Home | Online exhibits | The world's biomes

Marine regions cover about three-fourths of the Earth's surface and include oceans, coral reefs, and estuaries. Marine algae supply much of the world's oxygen supply and take in a huge amount of atmospheric carbon dioxide. The evaporation of the seawater provides rainwater for the land.

Oceans

The largest of all the ecosystems, oceans are very large bodies of water that dominate the Earth's surface. Like ponds and lakes, the ocean regions are separated into separate zones: intertidal, pelagic, abyssal, and benthic. All four zones have a great diversity of species. Some say that the ocean contains the richest diversity of species even though it contains fewer species than there are on land.

The intertidal zone is where the ocean meets the land — sometimes it is submerged and at other times exposed, as waves and tides come in and out. Because of this, the communities are constantly changing. On rocky coasts, the zone is stratified vertically. Where only the highest tides reach, there are only a few species of algae and mollusks. In those areas usually submerged during high tide, there is a more diverse array of algae and small animals, such as herbivorous snails, crabs, sea stars, and small fishes. At the bottom of the intertidal zone, which is only exposed during the lowest tides, many invertebrates, fishes, and seaweed can be found. The intertidal zone on sandier shores is not as stratified as in the rocky areas. Waves keep mud and sand constantly moving, thus very few algae and plants can establish themselves — the fauna include worms, clams, predatory crustaceans, crabs, and shorebirds.

The pelagic zone includes those waters further from the land, basically the open ocean. The pelagic zone is generally cold though it is hard to give a general temperature range since, just like ponds and lakes, there is thermal stratification with a constant mixing of warm and cold ocean currents. The flora in the pelagic zone include surface seaweeds. The fauna include many species of fish and some mammals, such as whales and dolphins. Many feed on the abundant plankton.

The benthic zone is the area below the pelagic zone, but does not include the very deepest parts of the ocean (see abyssal zone below). The bottom of the zone consists of sand, slit, and/or dead organisms. Here temperature decreases as depth increases toward the abyssal zone, since light cannot penetrate through the deeper water. Flora are represented primarily by seaweed while the fauna, since it is very nutrient-rich, include all sorts of bacteria, fungi, sponges, sea anemones, worms, sea stars, and fishes.

The deep ocean is the abyssal zone. The water in this region is very cold (around 3° C), highly pressured, high in oxygen content, but low in nutritional content. The abyssal zone supports many species of invertebrates and fishes. Mid-ocean ridges (spreading zones between tectonic plates), often with hydrothermal vents, are found in the abyssal zones along the ocean floors. Chemosynthetic bacteria thrive near these vents because of the large amounts of hydrogen sulfide and other minerals they emit. These bacteria are thus the start of the food web as they are eaten by invertebrates and fishes.

Coral reefs

Coral reefs are widely distributed in warm shallow waters. They can be found as barriers along continents (e.g., the Great Barrier Reef off Australia), fringing islands, and atolls. Naturally, the dominant organisms in coral reefs are corals. Corals are interesting since they consist of both algae (zooanthellae) and tissues of animal polyp. Since reef waters tend to be nutritionally poor, corals obtain nutrients through the algae via photosynthesis and also by extending tentacles to obtain plankton from the water. Besides corals, the fauna include several species of microorganisms, invertebrates, fishes, sea urchins, octopuses, and sea stars.

Estuaries

Estuaries are areas where freshwater streams or rivers merge with the ocean. This mixing of waters with such different salt concentrations creates a very interesting and unique ecosystem. Microflora like algae, and macroflora, such as seaweeds, marsh grasses, and mangrove trees (only in the tropics), can be found here. Estuaries support a diverse fauna, including a variety of worms, oysters, crabs, and waterfowl.

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Home | Online exhibits | The world's biomes Marine regions cover about three-fourths of the Earth's surface and include oceans, coral reefs, and estuaries. Marine algae supply much of the world's oxygen supply and take in a huge amount of atmospheric carbon dioxide. The evaporation of the seawater provides rainwater for the land.

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Home | Online exhibits | The world's biomes

Sand dunes in Death Valley National Monument, California.

Deserts cover about one fifth of the Earth's surface and occur where rainfall is less than 50 cm/year. Although most deserts, such as the Sahara of North Africa and the deserts of the southwestern U.S., Mexico, and Australia, occur at low latitudes, another kind of desert, cold deserts, occur in the basin and range area of Utah and Nevada and in parts of western Asia. Most deserts have a considerable amount of specialized vegetation, as well as specialized vertebrate and invertebrate animals. Soils often have abundant nutrients because they need only water to become very productive and have little or no organic matter. Disturbances are common in the form of occasional fires or cold weather, and sudden, infrequent, but intense rains that cause flooding.

There are relatively few large mammals in deserts because most are not capable of storing sufficient water and withstanding the heat. Deserts often provide little shelter from the sun for large animals. The dominant animals of warm deserts are nonmammalian vertebrates, such as reptiles. Mammals are usually small, like the kangaroo mice of North American deserts.

Desert biomes can be classified according to several characteristics.
There are four major types of deserts:

Hot and dry desert

The four major North American deserts of this type are the Chihuahuan, Sonoran, Mojave and Great Basin. Others outside the U.S. include the Southern Asian realm, Neotropical (South and Central America), Ethiopian (Africa) and Australian.

The seasons are generally warm throughout the year and very hot in the summer. The winters usually bring little rainfall.

From left: Baja, Mexico desert; desert in Uluru National Park, Australia; desert near the Kofa Mountains, Arizona.

Temperatures exhibit daily extremes because the atmosphere contains little humidity to block the Sun's rays. Desert surfaces receive a little more than twice the solar radiation received by humid regions and lose almost twice as much heat at night. Many mean annual temperatures range from 20-25° C. The extreme maximum ranges from 43.5-49° C. Minimum temperatures sometimes drop to -18° C.

Rainfall is usually very low and/or concentrated in short bursts between long rainless periods. Evaporation rates regularly exceed rainfall rates. Sometimes rain starts falling and evaporates before reaching the ground. Rainfall is lowest on the Atacama Desert of Chile, where it averages less than 1.5 cm. Some years are even rainless. Inland Sahara also receives less than 1.5 cm a year. Rainfall in American deserts is higher — almost 28 cm a year.

Soils are course-textured, shallow, rocky or gravely with good drainage and have no subsurface water. They are coarse because there is less chemical weathering. The finer dust and sand particles are blown elsewhere, leaving heavier pieces behind.

Canopy in most deserts is very rare. Plants are mainly ground-hugging shrubs and short woody trees. Leaves are "replete" (fully supported with nutrients) with water-conserving characteristics. They tend to be small, thick and covered with a thick cuticle (outer layer). In the cacti, the leaves are much-reduced (to spines) and photosynthetic activity is restricted to the stems. Some plants open their stomata (microscopic openings in the epidermis of leaves that allow for gas exchange) only at night when evaporation rates are lowest. These plants include: yuccas, ocotillo, turpentine bush, prickly pears, false mesquite, sotol, ephedras, agaves and brittlebush.

The animals include small nocturnal (active at night) carnivores. The dominant animals are burrowers and kangaroo rats. There are also insects, arachnids, reptiles and birds. The animals stay inactive in protected hideaways during the hot day and come out to forage at dusk, dawn or at night, when the desert is cooler.

Semiarid desert

The major deserts of this type include the sagebrush of Utah, Montana and Great Basin. They also include the Nearctic realm (North America, Newfoundland, Greenland, Russia, Europe and northern Asia).

The summers are moderately long and dry, and like hot deserts, the winters normally bring low concentrations of rainfall. Summer temperatures usually average between 21-27° C. It normally does not go above 38° C and evening temperatures are cool, at around 10° C. Cool nights help both plants and animals by reducing moisture loss from transpiration, sweating and breathing. Furthermore, condensation of dew caused by night cooling may equal or exceed the rainfall received by some deserts. As in the hot desert, rainfall is often very low and/or concentrated. The average rainfall ranges from 2-4 cm annually.

From left: Red Rock Canyon National Conservation Area, southern Nevada; sagebrush near Bridger, Montana; Castle Valley, Utah, east of Arches National Park.

The soil can range from sandy and fine-textured to loose rock fragments, gravel or sand. It has a fairly low salt concentration, compared to deserts which receive a lot of rain (acquiring higher salt concentrations as a result). In areas such as mountain slopes, the soil is shallow, rocky or gravely with good drainage. In the upper bajada (lower slopes) they are coarse-textured, rocky, well-drained and partly "laid by rock bench." In the lower bajada (bottom land) the soil is sandy and fine-textured, often with "caliche hardpan." In each case there is no subsurface water.

The spiny nature of many plants in semiarid deserts provides protection in a hazardous environment. The large numbers of spines shade the surface enough to significantly reduce transpiration. The same may be true of the hairs on the woolly desert plants. Many plants have silvery or glossy leaves, allowing them to reflect more radiant energy. These plants often have an unfavorable odor or taste. Semiarid plants include: Creosote bush, bur sage ( Franseria dumosa or F. deltoidea), white thorn, cat claw, mesquite, brittle bushes ( Encelia farinosa), lyciums, and jujube.

During the day, insects move around twigs to stay on the shady side; jack rabbits follow the moving shadow of a cactus or shrub. Naturally, many animals find protection in underground burrows where they are insulated from both heat and aridity. These animals include mammals such as the kangaroo rats, rabbits, and skunks; insects like grasshoppers and ants; reptiles are represented by lizards and snakes; and birds such as burrowing owls and the California thrasher.

Coastal desert

These deserts occur in moderately cool to warm areas such as the Nearctic and Neotropical realm. A good example is the Atacama of Chile.

The cool winters of coastal deserts are followed by moderately long, warm summers. The average summer temperature ranges from 13-24° C; winter temperatures are 5° C or below. The maximum annual temperature is about 35° C and the minimum is about -4° C. In Chile, the temperature ranges from -2 to 5° C in July and 21-25° C in January.

The average rainfall measures 8-13 cm in many areas. The maximum annual precipitation over a long period of years has been 37 cm with a minimum of 5 cm.

The soil is fine-textured with a moderate salt content. It is fairly porous with good drainage. Some plants have extensive root systems close to the surface where they can take advantage of any rain showers. All of the plants with thick and fleshy leaves or stems can take in large quantities of water when it is available and store it for future use. In some plants, the surfaces are corrugated with longitudinal ridges and grooves. When water is available, the stem swells so that the grooves are shallow and the ridges far apart. As the water is used, the stem shrinks so that the grooves are deep and ridges close together. The plants living in this type of desert include the salt bush, buckwheat bush, black bush, rice grass, little leaf horsebrush, black sage, and chrysothamnus.

Some animals have specialized adaptations for dealing with the desert heat and lack of water. Some toads seal themselves in burrows with gelatinous secretions and remain inactive for eight or nine months until a heavy rain occurs. Amphibians that pass through larval stages have accelerated life cycles, which improves their chances of reaching maturity before the waters evaporate. Some insects lay eggs that remain dormant until the environmental conditions are suitable for hatching. The fairy shrimps also lay dormant eggs. Other animals include: insects, mammals (coyote and badger), amphibians (toads), birds (great horned owl, golden eagle and the bald eagle), and reptiles (lizards and snakes).

Cold desert

These deserts are characterized by cold winters with snowfall and high overall rainfall throughout the winter and occasionally over the summer. They occur in the Antarctic, Greenland and the Nearctic realm. They have short, moist, and moderately warm summers with fairly long, cold winters. The mean winter temperature is between -2 to 4° C and the mean summer temperature is between 21-26° C.

The winters receive quite a bit of snow. The mean annual precipitation ranges from 15-26 cm. Annual precipitation has reached a maximum of 46 cm and a minimum of 9 cm. The heaviest rainfall of the spring is usually in April or May. In some areas, rainfall can be heavy in autumn. The soil is heavy, silty, and salty. It contains alluvial fans where soil is relatively porous and drainage is good so that most of the salt has been leached out.

The plants are widely scattered. In areas of shadscale, about 10 percent of the ground is covered, but in some areas of sagebush it approaches 85 percent. Plant heights vary between 15 cm and 122 cm. The main plants are deciduous, most having spiny leaves. Widely distributed animals are jack rabbits, kangaroo rats, kangaroo mice, pocket mice, grasshopper mice, and antelope ground squirrels. In areas like Utah, population density of these animals can range from 14-41 individuals per hectare. All except the jack rabbits are burrowers. The burrowing habit also applies to carnivores like the badger, kit fox, and coyote. Several lizards do some burrowing and moving of soil. Deer are found only in the winter.

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Home | Online exhibits | The world's biomes Sand dunes in Death Valley National Monument, California. Deserts cover about one fifth of the Earth's surface and occur where rainfall is less than 50 cm/year.

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Home | Online exhibits | The world's biomes

About 420 million years ago, during the Silurian Period, ancient plants and arthropods began to occupy the land. Over the millions of years that followed, these land colonizers developed and adapted to their new habitat. The first forests were dominated by giant horsetails, club mosses, and ferns that stood up to 40 feet tall.

Life on Earth continued to evolve, and in the late Paleozoic, gymnosperms appeared. By the Triassic Period (245-208 mya), gymnosperms dominated the Earth's forests. In the Cretaceous Period (144-65m mya), the first flowering plants (angiosperms) appeared. They evolved together with insects, birds, and mammals and radiated rapidly, dominating the landscape by the end of the Period. The landscape changed again during the Pleistocene Ice Ages — the surface of the planet that had been dominated by tropical forests for millions of years changed, and temperate forests spread in the Northern Hemisphere.

Today, forests occupy approximately one-third of Earth's land area, account for over two-thirds of the leaf area of land plants, and contain about 70% of carbon present in living things. They have been held in reverence in folklore and worshipped in ancient religions. However, forests are becoming major casualties of civilization as human populations have increased over the past several thousand years, bringing deforestation, pollution, and industrial usage problems to this important biome.

Present-day forest biomes, biological communities that are dominated by trees and other woody vegetation ( Spurr and Barnes 1980), can be classified according to numerous characteristics, with seasonality being the most widely used. Distinct forest types also occur within each of these broad groups.

There are three major types of forests, classed according to latitude:

Tropical forest

Left: Ranomafana National Park, Madagascar; right: Hawaiian forest.

Tropical forests are characterized by the greatest diversity of species. They occur near the equator, within the area bounded by latitudes 23.5 degrees N and 23.5 degrees S. One of the major characteristics of tropical forests is their distinct seasonality: winter is absent, and only two seasons are present (rainy and dry). The length of daylight is 12 hours and varies little.

Temperature is on average 20-25° C and varies little throughout the year: the average temperatures of the three warmest and three coldest months do not differ by more than 5 degrees.

Precipitation is evenly distributed throughout the year, with annual rainfall exceeding 200 cm.

Soil is nutrient-poor and acidic. Decomposition is rapid and soils are subject to heavy leaching.

Canopy in tropical forests is multilayered and continuous, allowing little light penetration.

Flora is highly diverse: one square kilometer may contain as many as 100 different tree species. Trees are 25-35 m tall, with buttressed trunks and shallow roots, mostly evergreen, with large dark green leaves. Plants such as orchids, bromeliads, vines (lianas), ferns, mosses, and palms are present in tropical forests.

Fauna include numerous birds, bats, small mammals, and insects.

Further subdivisions of this group are determined by seasonal distribution of rainfall:

evergreen rainforest: no dry season.

seasonal rainforest: short dry period in a very wet tropical region (the forest exhibits definite seasonal changes as trees undergo developmental changes simultaneously, but the general character of vegetation remains the same as in evergreen rainforests).

semievergreen forest: longer dry season (the upper tree story consists of deciduous trees, while the lower story is still evergreen).

moist/dry deciduous forest (monsoon): the length of the dry season increases further as rainfall decreases (all trees are deciduous).

More than one half of tropical forests have already been destroyed.

Temperate forest

Temperate forests occur in eastern North America, northeastern Asia, and western and central Europe. Well-defined seasons with a distinct winter characterize this forest biome. Moderate climate and a growing season of 140-200 days during 4-6 frost-free months distinguish temperate forests.

Temperature varies from -30° C to 30° C.

Precipitation (75-150 cm) is distributed evenly throughout the year.

Soil is fertile, enriched with decaying litter.

Canopy is moderately dense and allows light to penetrate, resulting in well-developed and richly diversified understory vegetation and stratification of animals.

Flora is characterized by 3-4 tree species per square kilometer. Trees are distinguished by broad leaves that are lost annually and include such species as oak, hickory, beech, hemlock, maple, basswood, cottonwood, elm, willow, and spring-flowering herbs.

Fauna is represented by squirrels, rabbits, skunks, birds, deer, mountain lion, bobcat, timber wolf, fox, and black bear.

Further subdivisions of this group are determined by seasonal distribution of rainfall:

moist conifer and evergreen broad-leaved forests: wet winters and dry summers (rainfall is concentrated in the winter months and winters are relatively mild).

dry conifer forests: dominate higher elevation zones; low precipitation.

mediterranean forests: precipitation is concentrated in winter, less than 100 cm per year.

temperate coniferous: mild winters, high annual precipitation (greater than 200 cm).

temperate broad-leaved rainforests: mild, frost-free winters, high precipitation (more than 150 cm) evenly distributed throughout the year.

Only scattered remnants of original temperate forests remain.

Boreal forest (taiga)

Boreal forests, or taiga, represent the largest terrestial biome. Occuring between 50 and 60 degrees north latitudes, boreal forests can be found in the broad belt of Eurasia and North America: two-thirds in Siberia with the rest in Scandinavia, Alaska, and Canada. Seasons are divided into short, moist, and moderately warm summers and long, cold, and dry winters. The length of the growing season in boreal forests is 130 days.

Temperatures are very low.

Precipitation is primarily in the form of snow, 40-100 cm annually.

Soil is thin, nutrient-poor, and acidic.

Canopy permits low light penetration, and as a result, understory is limited.

Flora consist mostly of cold-tolerant evergreen conifers with needle-like leaves, such as pine, fir, and spruce.

Fauna include woodpeckers, hawks, moose, bear, weasel, lynx, fox, wolf, deer, hares, chipmunks, shrews, and bats.

Current extensive logging in boreal forests may soon cause their disappearance.

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Home | Online exhibits | The world's biomes About 420 million years ago, during the Silurian Period, ancient plants and arthropods began to occupy the land. Over the millions of years that followed, these land colonizers developed and adapted to their new habitat.

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Home | Online exhibits | The world's biomes

Grasslands are characterized as lands dominated by grasses rather than large shrubs or trees. In the Miocene and Pliocene Epochs, which spanned a period of about 25 million years, mountains rose in western North America and created a continental climate favorable to grasslands. Ancient forests declined and grasslands became widespread. Following the Pleistocene Ice Ages, grasslands expanded in range as hotter and drier climates prevailed worldwide. There are two main divisions of grasslands:

Savanna

Savanna is grassland with scattered individual trees. Savannas of one sort or another cover almost half the surface of Africa (about five million square miles, generally central Africa) and large areas of Australia, South America, and India. Climate is the most important factor in creating a savanna. Savannas are always found in warm or hot climates where the annual rainfall is from about 50.8 to 127 cm (20-50 inches) per year. It is crucial that the rainfall is concentrated in six or eight months of the year, followed by a long period of drought when fires can occur. If the rain were well distributed throughout the year, many such areas would become tropical forest. Savannas which result from climatic conditions are called climatic savannas. Savannas that are caused by soil conditions and that are not entirely maintained by fire are called edaphic savannas. These can occur on hills or ridges where the soil is shallow, or in valleys where clay soils become waterlogged in wet weather. A third type of savanna, known as derived savanna, is the result of people clearing forest land for cultivation. Farmers fell a tract of forest, burn the dead trees, and plant crops in the ashes for as long as the soil remains fertile. Then, the field is abandoned and, although forest trees may recolonize, grass takes over on the bare ground (succession), becoming luxuriant enough to burn within a year or so. In Africa, a heavy concentration of elephants in protected parkland have created a savanna by eating leaves and twigs and breaking off the branches, smashing the trunks and stripping the bark of trees. Elephants can convert a dense woodland into an open grassland in a short period of time. Annual fires then maintain the area as a savanna.

The soil of the savanna is porous, with rapid drainage of water. It has only a thin layer of humus (the organic portion of the soil created by partial decomposition of plant or animal matter), which provides vegetation with nutrients. Savannas are sometimes classified as forests. The predominant vegetation consists of grasses and forbs (small broad-leaved plants that grow with grasses). Different savannas support different grasses due to disparities in rainfall and soil conditions. Because the savanna supports such a large number of species competing for living space, usually only one or a few kinds of grass are more successful than the others in a particular area. For example, in drier savannas such as those on the Serengeti plains or Kenya's Laikipia plateau, the dominant grasses on well-drained soils are Rhodes grass and red oat grass; throughout the East African savannas, star grasses are dominant; the lemon grasses are common in many western Uganda savannas. Deciduous trees and shrubs are scattered across the open landscape. One type of savanna common in southwestern Kenya, Tanzania, and Uganda, known as grouped-tree grassland, has trees growing only on termite mounds — the intervening soil being too thin or poorly drained to support the growth of trees at all. Frequent fires and large grazing mammals kill seedlings, thus keeping the density of trees and shrubs low. Savannas receive an average annual rainfall of 76.2-101.6 cm (30-40 inches).

Savanna has both a dry and a rainy season. Seasonal fires play a vital role in the savanna's biodiversity. In October, a series of violent thunderstorms, followed by a strong drying wind, signals the beginning of the dry season. Fire is prevalent around January, at the height of the dry season. Fires in savannas are often caused by poachers who want to clear away dead grass to make it easier to see their prey. The fires do not devastate the community. Most of the animals killed by the fires are insects with short life spans. A fire is a feast for some animals, such as birds that come to sites of fires to eat grasshoppers, stick insects, beetles, mice, and lizards that are killed or driven out by the fire. Underground holes and crevices provide a safe refuge for small creatures. Larger animals are usually able to run fast enough to escape the fire. Although the dry stems and leaves of grasses are consumed by fire, the grasses' deep roots remain unharmed. These roots, with all their starch reserves, are ready to send up new growth when the soil becomes more moist. The scattered shrubs can also subsist on food reserves in their roots while they await the time to venture above the soil again. Unlike grasses and shrubs, trees survive a fire by retaining some moisture in all their above-ground parts throughout the dry season. Sometimes they have a corky bark or semisucculent trunk covered with smooth resinous bark, both being fire resistant. A fire leaves scorched earth covered with a fine layer of powdery black ash in its wake. During March, violent thunderstorms occur again, this time heralding the rainy season. When the rains come, savanna bunch grasses grow vigorously. Some of the larger grasses grow an inch or more in 24 hours. The savannas experiences a surge of new life at this time. For example, many antelope calves are born. With so much grass to feed on, mothers have plenty of milk. Calves die if the rains fail to come.

Other animals (which do not all occur in the same savanna) include giraffes, zebras, buffaloes, kangaroos, mice, moles, gophers, ground squirrels, snakes, worms, termites, beetles, lions, leopards, hyenas, and elephants.

There are also some environmental concerns regarding savannas such as poaching, overgrazing, and clearing of the land for crops.

Temperate grassland

Temperate grasslands are characterized as having grasses as the dominant vegetation. Trees and large shrubs are absent. Temperatures vary more from summer to winter, and the amount of rainfall is less in temperate grasslands than in savannas. The major manifestations are the veldts of South Africa, the puszta of Hungary, the pampas of Argentina and Uruguay, the steppes of the former Soviet Union, and the plains and prairies of central North America. Temperate grasslands have hot summers and cold winters. Rainfall is moderate. The amount of annual rainfall influences the height of grassland vegetation, with taller grasses in wetter regions. As in the savanna, seasonal drought and occasional fires are very important to biodiversity. However, their effects aren't as dramatic in temperate grasslands as they are in savannas. The soil of the temperate grasslands is deep and dark, with fertile upper layers. It is nutrient-rich from the growth and decay of deep, many-branched grass roots. The rotted roots hold the soil together and provide a food source for living plants. Each different species of grass grows best in a particular grassland environment (determined by temperature, rainfall, and soil conditions). The seasonal drought, occasional fires, and grazing by large mammals all prevent woody shrubs and trees from invading and becoming established. However, a few trees, such as cottonwoods, oaks, and willows grow in river valleys, and some nonwoody plants, specifically a few hundred species of flowers, grow among the grasses. The various species of grasses include purple needlegrass, blue grama, buffalo grass, and galleta. Flowers include asters, blazing stars, coneflowers, goldenrods, sunflowers, clovers, psoraleas, and wild indigos.

Precipitation in the temperate grasslands usually occurs in the late spring and early summer. The annual average is about 50.8 to 88.9 cm (20-35 inches). The temperature range is very large over the course of the year. Summer temperatures can be well over 38° C (100 degrees Fahrenheit), while winter temperatures can be as low as -40° C (-40 degrees Fahrenheit).

The fauna (which do not all occur in the same temperate grassland) include gazelles, zebras, rhinoceroses, wild horses, lions, wolves, prairie dogs, jack rabbits, deer, mice, coyotes, foxes, skunks, badgers, blackbirds, grouses, meadowlarks, quails, sparrows, hawks, owls, snakes, grasshoppers, leafhoppers, and spiders.

There are also environmental concerns regarding the temperate grasslands. Few natural prairie regions remain because most have been turned into farms or grazing land. This is because they are flat, treeless, covered with grass, and have rich soil.

Temperate grasslands can be further subdivided. Prairies are grasslands with tall grasses while steppes are grasslands with short grasses. Prairie and steppes are somewhat similar but the information given above pertains specifically to prairies — the following is a specific description of steppes.

Steppes are dry areas of grassland with hot summers and cold winters. They receive 25.4-50.8 cm (10-20 inches) of rainfall a year. Steppes occur in the interiors of North America and Europe. Plants growing in steppes are usually greater than 1 foot tall. They include blue grama and buffalo grass, cacti, sagebrush, speargrass, and small relatives of the sunflower. Steppe fauna includes badgers, hawks, owls, and snakes. Today, people use steppes to graze livestock and to grow wheat and other crops. Overgrazing, plowing, and excess salts left behind by irrigation waters have harmed some steppes. Strong winds blow loose soil from the ground after plowing, especially during droughts. This causes the dust storms of the Great Plains of the U.S.

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Home | Online exhibits | The world's biomes Grasslands are characterized as lands dominated by grasses rather than large shrubs or trees. In the Miocene and Pliocene Epochs, which spanned a period of about 25 million years, mountains rose in western North America and created a continental climate favorable to grasslands.

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