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He mated male and female fruit flies, by choosing three recessive traits: yellow body, miniature wings and white eyes (heterozygous female ) to a non a fly that had dominant traits of a brown boded, red eyed, large winged male. he concluded that the phenotype of male hybrids is contributed to the mother's X chromosome. Half the flies were dominant and the other half were resseive.

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~ Drew the outline of the first gene map of an X chromosome
~ Based it on the work of the Belgian cytologist F. A Janssens
~ Crossing over was another major influence on his gene map
~ Discovered rare cross overs which are also called double cross overs
~ Did all his work while he was still a student, he completed it as part of his PHD thesis